Prostaglandins, Leukotrienesand Essential Fatfy Acids (1998) 58(6), 421424
© HarcourtBrace& Co. Ltd 1998
Effects of lysine clonixinate on
cyclooxygenase I and II in rat lung
and stomach preparations
A. M. Franchi, G. Di Girolamo, A. R. de los Santos, M. L. Marti, M. A. F. Gimeno
Centro de Estudios Farmacologicos y Botanicos (CEFYBO), Consejo Nacional de Investigaciones Cientfficas y Tecnicas (CONICET), Serrano 669, (1414)
Buenos Aires, Argentina
Summary Lysine clonixinate (LC) is a drug of antiinflammatory antipyretic and analgesic activity that produces minor
digestive side-effects. This fact induced us to think that LC is possibly a weak COX-1 inhibitor. In order to investigate
our hypothesis we inhibited cyclooxygenase activity with LC or indomethacin (INDO) in rat lung and stomach obtained
from rats treated with lipopolysacharide (LPS) and control rats. Rat lung preparations incubated with 14C-arachidonic
acid synthesise mainly PGE2. LC at 2.5 and 4.1 x 104 M does not modify the basal production of PGE 2 (probably
COX-l) but at 6.8 x 10-2 M significantly inhibited PGE2 production (approximately 48.5% inhibition, P < 0.001). On the
other hand, INDO at 104 inhibited the basal production of PGE 2 by around 73%. In LPS-treated rats, the production of
PGE2 was significantly higher than in the lungs of control rats, probably due to the induction of COX-2. The addition of
LC at 2.7 and 4.1 x 10-2 M recovered the control values of PGE2 inhibiting, probably only from COX-2 activity. LC at
higher concentrations (6.8 x 10-2 M) and INDO 104 M inhibited PGE2 formed by COX-2 and also partly by COX-1
activity.
INTRODUCTION
The PGH2 synthase is a rate-limiting enzyme in prosta-
glandin and thromboxane synthesis, it possesses two
enzymatic activities, a cyclooxygenase which catalyses
the oxygenation of arachidonic acid to PGG2 and per-
oxidase which reduces PGG2 to PGH2.~ Until recently, a
single cyclooxygenase (COX) was recognized as the
enzyme responsible for the generation of the prosta-
glandins. However, a second form of the enzyme has now
been identified, which is induced by proinflammatory
cytokines, growth factor hormones and oncogenes. This
cyclooxygenase (COX-2) is specifically associated with
cells and tissues involved in inflammation.2-4 Actually,
cyclooxygenase has been found in two forms, namely the
constitutive, widely occurring enzyme (COX-l) and the
inducible isoform (COX-2), which is expressed after stim-
ulation with endotoxin lipopolysacharide (LPS) or some
cytokines? The LPS of gram-negative bacteria elicit a wide
range of effects in biological systems in vivo and in vitro.
Received 8 November 1996
Accepted 26 May 1998
Correspondence to: A. M. Franchi
Several cell types and tissues, such as lung challenged
with LPS, produce increased amounts of both nitric oxide 5
and prostaglandins. 6 Systemic administration of LPS
resulted in induction of COX-2 mRNA within 1 h and
inducible nitric oxide synthase (iNOS) in lung, without
affecting levels of COX-1. 7 Previous studies in vitro sug-
gested that side-effects of non-steroidal antfinflammatory
drugs (NSMDS) correlate with their ability to inhibit COX-
1, while the antiinflammatory effects are due to their
ability to inhibit COX-2. On the other hand, the anti-
inflammatory steroid dexamethasone completely inhibited
the expression of COX-2, whereas COX-1 was unaffected
in macrophages stimulated with LPSfi
The unique involvement of the isoform COX-2 in
inflammatory disorders has provided an exciting opportu-
nity for developing pharmacologic agents that selectively
inhibit COX-2. Most of the currently available NSMDs are
generally one or two orders of magnitude more potent in
bloclrang COX-1.9 Lysine clonixinate (LC) is a drug of anti-
inflammatory, antipyretic and analgesic activity which
has proven to be rapid and effective in clinical uses. ~°-12 It
has been reported that LC produces minor digestive side-
effects. ~3 This fact induces us to think that this drug is
possibly a weak COX-1 inhibitor; in the rat lung a high
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