Aqueous Two-Phase System Formation Kinetics for Elastin-Like
Polypeptides of Varying Chain Length
Yanjie Zhang, Kimberly Trabbic-Carlson, Fernando Albertorio, Ashutosh Chilkoti, and
Paul S. Cremer*
Department of Chemistry, Texas A&M University, 3255 TAMU, College Station, Texas 77843, and
Department of Biomedical Engineering, Duke University, Durham, North Carolina 27708
Received March 18, 2006; Revised Manuscript Received May 1, 2006
The kinetics of aqueous two-phase system (ATPS) formation for elastin-like polypeptides (ELP) with defined
chemical composition and chain length was investigated by dark field microscopy in an on-chip format with a
linear temperature gradient. Scattering intensities from peptide solutions in the presence and absence of sodium
dodecyl sulfate (SDS) were recorded as a function of temperature and time, simultaneously. It was found that the
formation of the ATPS for three ELPs of different molecular weights (36 075, 59 422, and 129 856 Da) in the
absence of SDS followed a coalescence mechanism, and the rate constant and activation energy were independent
of chain length. With the introduction of SDS into the ELP solutions, the rate constants were attenuated more
strongly with increasing chain length. Moreover, the coalescence process in the presence of SDS showed non-
Arrhenius kinetics as a function of temperature. For the two shorter ELPs, ATPS formation occurred via coalescence
at all SDS concentrations and temperatures investigated. On the other hand, the coalescence process was greatly
suppressed for the longest ELP at elevated temperatures and higher SDS concentrations. Under these circumstances,
ATPS formation was forced to proceed via a mixed Ostwald ripening and coalescence mechanism.
Introduction
The formation of an aqueous two-phase system
1-3
com-
mences when a macromolecular system spontaneously separates
into two separate water-containing phases. Such phenomena
often involve two polymers separating into distinct phases with
different densities. ATPS formation may also occur in solutions
consisting of a single polymer plus a high concentration of an
appropriate salt
4-7
or even in the absence of salt for certain
thermoresponsive polymers.
8-10
In this final case, ATPS forma-
tion occurs above the polymer’s lower critical solution temper-
ature (LCST). For these systems, a denser polymer-rich phase
is formed on the bottom of the container, while the upper phase
consists primarily of pure water. Since the phase boundary in
an ATPS has very low interfacial tension,
11
these systems have
been widely applied as tools for bioseparations without signifi-
cant interfacial denaturing effects.
12-14
Despite the practical and fundamental significance of ATPS
formation, there is almost no literature dealing with its formation
kinetics because of the difficulties involved in data collection.
To remedy this problem, we recently developed a temperature-
gradient microfluidic technique
15-17
that allows ATPS formation
to be followed in a high-throughput, low sample volume
fashion.
18
This enabled us to explore the activation energy of
the process for R-elastin, a thermoresponsive protein, which was
prepared by the methods of Partridge et al.
19
It was found that
there are two major mechanistic pathways through which ATPS
formation could proceed: coalescence and Ostwald ripening.
Coalescence is the direct fusion of one particle with another to
make a larger one, while Ostwald ripening involves the growth
of particles through the transfer of individual molecules between
them such that bigger particles grow at the expense of smaller
ones.
20
Each process has a different activation barrier associated
with it. In the case of coalescence, the barrier comes from the
removal of intervening solvent molecules between the particles
which are being joined. On the other hand, Ostwald ripening
involves the desorption of single macromolecules from shrinking
particles and their deposition onto growing particles. The rate
constants of ATPS formation for elastin are consistent with a
coalescence mechanism, although the process can be forced to
proceed through Ostwald ripening by the addition of sodium
dodecyl sulfate to the solution.
We suspected that the mechanism of ATPS formation might
be dependent on the molecular weight of the polymer; however,
the natural product, R-elastin, is a mixture of cross-linked
materials that is highly polydisperse. Therefore, to obtain more
control over the molecular weight, we employed elastin-like
polypeptides. ELPs are based on a repetitive pentapeptide motif,
Val-Pro-Gly-Xaa-Gly, where the guest residue, Xaa, is any
amino acid except Pro. The sequence and chain length can be
precisely controlled by genetic expression in bacteria using
recombinant DNA techniques.
21,22
The work described herein
was undertaken with three different molecular weight species
(36 075, 59 422, and 129 856 Da) to investigate the effect of
chain length on the kinetics of ATPS formation. It was found
that coalescence in the absence of SDS is not dependent on the
chain length of the ELPs; however, SDS increases the activation
barrier for the coalescence process of longer-chain ELPs to a
greater extent than for shorter ones. In fact, the ATPS formation
process could be forced to go by Ostwald ripening at higher
temperatures and SDS concentrations in the case of the 129 856
Da ELP, but not for the two smaller molecules.
Experimental Section
The ELPs employed herein come from a library of ELP[V5A2G3-n]
where the repeat sequence of amino acids is the same. Members within
the library differ only in their chain length, where n is the number of
pentapeptides. For example, ELP[V5A2G3-90] consists of 90 pentapep-
tides and has a repeat unit composed of 10 pentapeptides with the guest
2192 Biomacromolecules 2006, 7, 2192-2199
10.1021/bm060254y CCC: $33.50 © 2006 American Chemical Society
Published on Web 06/20/2006