- Variation in modern pollen from tropical evergreen forests and the monsoon seasonality gradient - 551
Journal of Vegetation Science 14: 551-562, 2003
© IAVS; Opulus Press Uppsala.
Abstract. This study analyses the pollen signature of tropical
lowland forests (< 1000 m a.s.l.) in the Asian monsoon cli-
mate. Its aim is to investigate how well the pollen data can
reproduce the vegetation patterns in tropical India, and how
the variations in the pollen composition are related to the
gradient of decreasing plant moisture availability (measured
by the ratio of actual over equilibrium evapotranspiration) that
is associated with the strong seasonality of precipitation that
characterizes the monsoon climate regime.
We used canonical correspondence analysis (CCA) to
relate the variations in the pollen composition of 71 surface
soil samples from evergreen and semi-evergreen forests dis-
tributed along the western coast of south India (8° 48' N-15°
08' N), with the climate characteristics of the sampling sites.
We show that variations in plant moisture availability strongly
determine variations in the pollen composition; for example
evergreen and semi-evergreen forests can be distinguished on
the basis of their pollen assemblages. Variations in the mean
temperature of the coldest month associated with elevation
also determine distinct pollen assemblages; for example ever-
green forests above 800 m a.s.l. present different pollen signa-
tures than those below this altitude/temperature limit. Varia-
tions in the relative abundance of some pollen taxa are strongly
related to plant moisture availability and taxa indicators of
climate can be identified. Hence, modern pollen assemblages
from tropical forests in south India carry considerable infor-
mation about vegetation patterns and climate. Paleoclimatic
changes, notably in the monsoon season, could be quantified.
Keywords: CCA; Dry season; Indicator taxa; Western Ghats.
Abbreviations: IVI = Importance value index; PANN = Mean
annual rainfall; MTCO = Mean temperature of coldest month.
Introduction
Pollen data from dated sedimentary sequences have
been widely used to reconstruct vegetation patterns
during past periods (e.g. Prentice & Webb 1998) and to
infer the changes in climate that have led to the observed
changes in vegetation distribution (Howe & Webb 1983;
Bonnefille et al. 1990; Cheddadi et al. 1997; Bonnefille
& Chalié 2000). Modern pollen surface samples from
known environments provide an important way to es-
tablish whether specific types of vegetation have a suf-
ficiently distinctive pollen spectrum to allow their iden-
tification in the fossil record despite inter-species differ-
ences in pollen production or post-depositional modifi-
cation. These investigations can also help to isolate the
bioclimate parameter(s) determining the limits of veg-
etation types (Bush 2000). These relationships then can
be used to improve inferences about past climates using
pollen-based reconstructions of vegetation patterns.
Recent investigations of modern pollen surface sam-
ples from the Indian subcontinent have shown that tem-
perature controls vegetation types along altitudinal gra-
dients (Bonnefille et al. 1999). Precipitation-related dif-
ferences between evergreen and deciduous forests also
can be distinguished on the basis of their pollen spectra
(Anupama et al. 2000; Barboni & Bonnefille 2001).
However, seasonality of precipitation, and specifically
the length of the dry season, may be a more important
determinant of the distribution of evergreen forest types
in southern India than precipitation (Pascal 1988). In
this paper we present analyses of modern pollen surface
samples from the evergreen forests in this region. Our
aims are to establish whether different floristic types of
evergreen vegetation can be distinguished by their pol-
len assemblages and to investigate the bioclimatic con-
trols on evergreen vegetation types.
Study area
This study concerns different forest types located in
the western Ghats, a narrow mountainous chain along
the west coast of south India. We focused on the area
between 8° and 15° N, well studied by botanists and
ecologists who provided maps of vegetation and bioclimate.
We restricted our study to areas below 1000 m a.s.l.
Variation in modern pollen from tropical evergreen forests
and the monsoon seasonality gradient in SW India
Barboni, D.
1, 2*
; Bonnefille, R.
2
; Prasad, S.
1
& Ramesh, B.R.
1
1
French Institute of Pondicherry, 11 Saint Louis Street, B.P.33, Pondicherry, 605001 India;
2
C.E.R.E.G.E., Europôle Méditerranéen de l’Arbois, B.P. 80, F-13545 Aix-en-Provence Cedex 4, France;
*
Corresponding author; E-mail barboni@cerege.fr