Animal Reproduction Science, 23 ( 1990 ) 213-222 213
Elsevier Science Publishers B.V., Amsterdam
Proteolytic and antiproteolytic activities in goat
antral follicles
A.M. Rosales b, O. Hernandez-Perez a, R. Dominguez a, E. Mercado a and
A. Rosado c
aLaboratorio de Biologia Molecular, Unidad de Investtgacion Biomedica, I.M.S.S., Mexico,
D.F. 03020, Mexico
bDepartamento de Produccion Agricola y Animal, Universidad Autonoma Metropolitana, Unidad
Xochimilco, Calzada del Hueso 1100, Mexico, D.F. 0496 O, Mexico (for correspondence )
cUniversidad Autonoma Metropolitana, Unidad lztapalapa, Mexico, D.F., Mexico
(Accepted 20 February 1990)
ABSTRACT
Rosales, A.M., Hernandez-Perez, O., Dominguez, R., Mercado, E. and Rosado, A., 1990. Proteolytic
and antiproteolytic activities in goat antral follicles. Anita. Reprod. Sci., 23: 213-222.
Goat antral follicles were dissected and classified initially by size into three groups: < 3 mm, be-
tween 3 and 6 ram, and > 6 ram. Follicles less than 3 mm diameter were studied without any further
classification. All others were classified according to their gross stereoscopic microscope appearance
into four sub-groups. Histological observation showed that sub-group 1 consisted mainly of non-atretic
follicles, sub-group 2 of follicles in the early stages of atresia, while sub-groups 3a and 3b were tbrmed
by secondary and tertiary atretic follicles. Tryptic and anti-trypic activites were measured in follicular
fluid, homogenized follicular cells and empty follicular sacs, using a fluorometric procedure. Anti-
tryptic activity in follicular fluid was found to be directly correlated with progress of atresia and in-
versely with follicular growth, being smallest in big ( > 6 mm), non-atretic follicles and consistently
higher in small ( < 3 mm) and in atretic follicles. No differences were detected between sub-group 3
follicles of different sizes. After elimination of anti-trypic activity by gel filtration, trypsin-like activity
in cell-free follicular fluid was found only on > 6 mm non-atretic follicles. High trypsin-like activity
was found in the cell walls of non-atretic or early atretic follicles. This activity was significantly lower
in sub-group 3a follicles and was almost absent in tertiary atretic follicles (sub-group 3b). In contrast,
trypsin-like activity in granulosa cells recovered by centrifugation of follicular fluid was twice as high
in sub-group 3a as in sub-groups 1 and 2 and almost five times higher than in sub-group 3b.
INTRODUCTION
Changes in follicular fluid steroid concentrations have been consistently
used as predictors of follicular maturation and atresia (Moor et al., 1978;
Yoshimura and Wallach, 1987). However, steroid hormones are, generally,
only intermediaries in the regulation of basic biochemical processes. There-
fore it would be of major interest to study other biochemical parameters of
0378-4320/90/$03.50 © 1990 -- Elsevier Science Publishers B.V.