Pharmacology Biochemistry & Behavior, Vol. 17, pp. 367-369, 1982. Printed in the U.S.A.
Tofizopam Modulates the Affinity of
Benzodiazepine Receptors in the Rat Brain
VEIJO SAANO AND ARTO URTTI*
Department of Pharmacology and Toxicology
and *Department of Pharmacy, University of Kuopio, P.O. Box 138, 70101 Kuopio 10, Finland
Received 19 October 1981
SAANO, V. AND A. URTTI. Tofizopam modulates the affinity ofbenzodiazepine receptors in the rat brain. PHARMAC.
BIOCHEM. BEHAV. 17(2) 367-369, 1982.--Tofizopam, a 3,4-benzodiazepine, lacks the sedative action common to
1,4-benzodiazepines, but has anxiolytic activity. In this study we administered tofizopam (50 mg/kg) to rats perorally twice
a day for six days, and analyzed the binding of [SH]flunitrazepam to benzodiazepine receptors of these drug-treated rats.
The effect of tofizopam treatment was compared to that brought about by treatment with diazepam (12 mg/kg twice a day
for six days) and to binding in controls treated with vehicle. Compared to the controls, the diazepam group had a marked
decrease in binding of [3H]flunitrazepam to benzodiazepine receptors both in the forebrain and in the hindbrain. As a result of
the increased affinity of the receptors tofizopam slightly, but statistically significantly, enhanced binding. With both drugs
the number of receptors was unaltered. The effect of tofizopam in the hindbrain was similar to that in the forebrain. The
results of this study support our earlier finding from single-dose studies that tofizopam acts indirectly on benzodiazepine
receptors.
Tofizopam Diazepam Benzodiazepine receptors
TOFIZOPAM, a 3,4-benzodiazepine, has been shown to
have an anxiolytic effect [4, 10, 17]. In contrast to 1,4-ben-
zodiazepines (e.g., diazepam, flunitrazepam, etc.), to-
fizopam does not cause sedation or muscle relaxation [12,
15, 17]. The therapeutic potency of various 1,4-ben-
zodiazepines correlates positively with their ability to
displace radiolabeled benzodiazepines from benzodiazepine
receptors in the central nervous system [8]. Tofizopam does
not displace tritiated flunitrazepam from receptors, but in
vitro enhances the receptor-specific binding of flunitrazepam
to rat brain and in vivo has been found to enhance this bind-
ing in rats after acute treatment [14]. The aim of the present
study was to analyze the effects of tofizopam on benzodiaz-
epine receptors after prolonged treatment. A small group of
rats was treated with diazepam so that we could compare the
effects of tofizopam to those caused by a 1,4-benzo-
diazepine; diazepam has been shown to occupy benzodiaz-
epine receptors during acute and chronic treatment to the
extent that is easy to identify in vitro using homogenates of
brain tissue from drug-treated animals [6,11].
METHOD
Twenty-three male rats of inbred strain (BD IX/Kuo)
weighing 220-340 g were housed in opaque plastic boxes
under standard laboratory conditions: 10 hr dark/14 hr light
cycle, air temperature 20-+0.5°C, relative air humidity 55-
75 ° . Before and during drug treatment, animals had free ac-
cess to food and tap water.
Tofizopam and diazepam were suspended in 1% solution
of Tween 80/water and were administered through a stomach
tube in a volume of 5 ml/kg body weight. Drugs (or vehicle to
controls) were administered twice a day at 12 hr intervals.
Each dose of tofizopam was 50 mg/kg; each dose of di-
azepam was 12 mg/kg. [3H]flunitrazepam was used as a
ligand; it had a specific activity of 86.4 Ci/mmole and was
purchased from New England Nuclear, Boston, MA. To-
fizopam, diazepam, and non-radioactive flunitrazepam were
generous gifts from the Research Center of Farmos Group,
Turku, Finland.
Drug treatment continued for six days. Rats were decapi-
tated 2 hr after they had received the last dose of drug. After
decapitation the brain was removed immediately and the
forebrain was separated from the hindbrain by an incision
posterior to the occipital lobes and across the brain stem.
The hindbrain (cerebellum + pons and medullar oblongata)
and the forebrain were homogenized individually in 32 vol-
umes of ice-cold Tris-HC1 buffer (50 mM, pH 7.4). Aliquots
of homogenate (400/zl) were added to a set of duplicate test
tubes containing 6-9 concentrations of [aH[flunitrazepam
(range from 0.06 to 8.12 nM). For determination of
nonspecific binding another set of tubes also contained 10
/zM concentration of non-radioactive flunitrazepam. The
tubes were incubated at 4°C for 30 minutes before the con-
tents were filtered through Whatman GF/B filters which
were then placed in 6 ml of Aquasol ® scintillation liquid for
at least 16 hr, after which the radioactivity was measured.
The protein content of each homogenate was determined
using the method described by Lowry et al. [7]. Maximum
binding and the dissociation constant for [3H]flunitrazepam
binding were determined by Scatchard analysis as described
by Bennett [2]. Statistical significance of the differences be-
tween values from drug-treated rats and those from control
rats was tested using Student's t-test.
Copyright © 1982 ANKHO International Inc.--0091-3057/82/080367-03503.00/0