Pharmacology Biochemistry & Behavior, Vol. 25, pp. 693-695, 1986.©AnkhoInternational Inc. Printed in the U.S.A. 0091-3057/86$3.00 + .00
BRIEF COMMUNICATION
The Effects of Apomorphine on Leverpress
Shock Escape Learning in Rats'
BRUCE A. MATTINGLY
Department of Psychology, Morehead State University, Morehead, KY40351
Received 24 March 1986
MATTINGLY, B. A. The effects of apomorphine on leverpress shock escape learning in rats. PHARMACOL BIOCHEM
BEHAV 25(3)693-695, 1986.--Four groups of rats (n = l0 each) were tested on a discrete trial leverpress shock escape task
15 rain following an intraperitoneal injection of either 0 (saline), 0.5, 1.0, or 2.0 mg/kg apomorphine hydrochloride. The
results indicated that all doses of apomorphine produced a severe disruption in escape performance. This disruption was
temporary, however, as all apomorphine groups were responding as quickly as the saline control rats by the end of the
training session. A comparison of the effects of apomorphine with the previously reported effects of scopolamine and septal
lesions on shock escape learning revealed both similarities and differences. These findings suggest that a septal lesion-
induced reduction of acetylcholine levels does not simply "unleash" an antagonistic dopaminergic system.
Rats Shock escape learning Apomorphine Dopaminergic mechanisms Scopolamine
Cholinergic mechanisms Septal lesions Acetylcholine Dopamine Acetylcholine-dopamine balance
FOLLOWING septal damage there is a significant reduction
in the levels of several forebrain neurotransmitters (see [3,9]).
Over the past few years we have been studying the potential
involvement of these secondary neurochemical changes in
the behavioral effects of septal lesions in certain aversive
learning tasks [4--7]. We have found, for example, that the
cholinergic antagonist, scopolamine, but not methyl-
scopolamine, produces a disruption in discrete trial lever-
press shock escape learning very similar to that produced by
septal lesions [4]. In contrast, a depletion of brain serotonin
produced by para-chlorophenylalanine (PCPA) does not af-
fect the performance of rats on a leverpress shock escape
task [6]. Likewise, increasing brain serotonin levels via
5-hydroxytryptophan does not affect the shock escape per-
formance of either normal or septally-lesioned rats [6].
Hence, these findings suggest that a lesion-induced reduc-
tion in brain acetylcholine, but not serotonin, levels may be
involved in the deficient shock escape performance of rats
with septal lesions.
Although our previous studies suggest the involvement of
cholinergic mechanisms, because of the interaction among
various neurotransmitters in many central "pathways,"
many neurotransmitters probably play some functional role
in any complex response pattern of the organism. For
example, in some areas of the brain (e.g., striatum), acetyl-
choline and dopamine have been found to function in precise
balance such that decreasing the activity of one has more or
less the same effect as mcreasing the activity of the other
(see [1] for review). Moreover, recent evidence has been
presented supporting a cholinergic-dopaminergic interaction
in various limbic structures including the septum (e.g., [8]).
It is possible, therefore, that the disruption of shock escape
learning produced by scopolamine and septal lesions may be
the result of an imbalance between acetylcholine-dopamine
mechanisms. If so, then increasing dopaminergic activity
should have an effect on leverpress shock escape perform-
ance in rats similar to that produced by decreasing acetyl-
choline activity with scopolamine or septal lesions. The pur-
pose of the present study, therefore, was to determine
whether the leverpress shock escape performance of rats
would be disrupted by a drug-induced facilitation of
dopaminergic activity. Consequently, groups of rats were
injected with either saline (control) or the central
dopaminergic agonist, apomorphine (0.5, 1.0, or 2.0 mg/kg)
and then tested on a discrete trial leverpress shock escape
task.
METHOD
Subjects
Forty male Wistar albino rats were experimentally naive
and approximately 90 days old on the day of testing. All rats
were housed individually and maintained on ad lib food and
water. A 12 hour light-dark cycle was held constant through-
out the experiment.
1This research was supported by a Faculty Research Grant from Morehead State University. The author is grateful to June Spencer and
Carlos Matin for their assistance in behavioral testing.
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