Research Article
Received: 3 October 2007, Revised: 26 August 2008, Accepted: 2 September 2008 Published online 31 October 2008 in Wiley Interscience
(www.interscience.wiley.com) DOI 10.1002/pca.1102
Copyright © 2008 John Wiley & Sons, Ltd. Phytochem. Anal. 2009; 20: 98–103
98
John Wiley & Sons, Ltd.
Polyphenolic composition of roots and
petioles of Rheum rhaponticum L.
Tõnu Püssa,
a
Piret Raudsepp,
a
Kristina Kuzina
b
and Ain Raal
b
*
ABSTRACT:
Introduction – Various species of the genus Rheum (Polygonaceae) are known for their high content of medicinally important
hydroxyanthraquinones. However, little information is available concerning the polyphenolic composition of garden or
dietary rhubarb Rheum rhaponticum L. (R. rhaponticum).
Objective – Determination of further polyphenols in the roots and petioles of R. rhaponticum.
Methodology – The dried plant material was extracted with 10-fold excess (v/w) of methanol and subsequently diluted five times
with methanol–water (1:1) and analysed by reversed-phase liquid chromatography using tandem UV-photodiode array and mass
selective detection (RP-HPLC-UV-ESI/MS
2
). Polyphenols were identified using either HPLC-ESI/MS
2
data obtained for respective
commercial standards or by comparison of a parent ion fragmentation picture with the respective MS
2
spectrum from the literature.
Results – The roots of R. rhaponticum were very rich in various hydroxystilbenes and contained four main substance groups –
derivatives of trans-piceatannol, trans-resveratrol, trans-rhapontigenin and trans-deoxyrhapontigenin. Additionally, pteros-
tilbene acetylglucosides and a number of hydroxyanthraquinones and their glycosides were identified in the root samples.
The profile of polyphenols in the petioles of R. rhaponticum was similar to that of the roots but the content of individual
substances was remarkably lower. The petioles of the R. rhaponticum additionally contained significant amounts of derivatives
of flavonol quercetin, which is a good antioxidant.
Conclusion – The study has shown that roots of R. rhaponticum contain a wide variety of hydroxystilbenes and deserve further
consideration as a source of medicinally interesting compounds. Copyright © 2008 John Wiley & Sons, Ltd.
Keywords: HPLC-UV-ESI/MS
2
; polyphenols; hydroxystilbenes; hydroxyanthraquinones; Rheum rhaponticum; garden rhubarb
Introduction
Different species of the genus Rheum (Polygonaceae), such as
R. palmatum L., R. emodi Wall. ex Meisn. and R. officinale Baill. are
known for their high content of physiologically active hydroxy-
anthraquinones (aloe-emodin, emodin, chrysophanol, rhein,
physcion and their glycosides) in their roots (Okabe et al., 1973;
Agarwal et al., 2000, Komatsu et al., 2006). On the other hand,
a number of hydroxystilbenes and their derivatives have been
identified in the rhizome of Korean rhubarb R. undulatum L.
(Kashiwada et al., 1984; Ko, 2000; Matsuda et al., 2000a,b;
Kageura et al., 2001), in the roots of R. maximowiczii Loinsk
(Shikishima et al., 2001) and in the rhizome of R. emodi (Babu et
al., 2004). More than 100 different phenolic compounds have
either been identified or tentatively characterised in six rhubarb
species (R. officinale, R. palmatum, R. tanguticum, R. franzenbachii,
R. hotaoense and R. emodi) by their parent ion fragmentation
spectra by Ye and co-authors (Ye et al., 2007).
There is substantially less data in the literature on the content
of phenolic compounds in the roots and especially in petioles of
garden or dietary rhubarb R. rhaponticum L., which in many
aspects is similar to R. undulatum. R. rhaponticum, the petioles of
which are used as a food component, has been known in Chinese
ethnopharmacology from ancient times. It has been shown that
the rhizomes of this rhubarb contain hydroxystilbene rhaponticin
(synonyms rhapontin and rhapontigenin glucoside), as well as
its precursor trans-resveratrol (Rupprich et al., 1980). The latter is
widely known for its versatile therapeutic potential (Piver et al.,
2003; Baur and Sinclair, 2006). The physiological properties of
resveratrol oligomers and related hydroxystilbenes have been
much less studied. Resveratrol oligomers usually support the
action of the monomer, but they may act also in a different manner
(Piver et al., 2003). For example, it has been demonstrated that
resveratrol oligomers strongly suppress HL-60 cell proliferation
and induce DNA damage (Kang et al., 2003). Rhaponticin or
rhapontin elicits antithrombotic and antiallergic properties (Park
et al., 2002) and isorhapontigenin attenuates cardiac hypertrophy
(Li et al., 2005).
A special extract from the roots of R. rhaponticum, referred to
as ERr 731® (trade name Phyto-Strol®, Carl Müller, Göppingen,
Germany), has been regularly prescribed for climacteric com-
plaints since 1993, and has been used in Germany for decades
for women of child-bearing age suffering from oligomenorrhoea
or amenorrhoea. According to recent investigations, the main
active components in this extract are hydroxystilbenes (Möller
et al., 2007; Wober et al., 2007).
A limited number of references can be found concerning
the determination of complex glycosides such as O-acetyl- or
O-galloylglucosides of hydroxystilbenes or hydroxyanthraquinones
in plants (Okasaka et al., 2004; Komatsu et al., 2006; Ye et al.,
* Correspondence to: A. Raal, Institute of Pharmacy, University of Tartu,
Nooruse 1, 50411 Tartu, Estonia. E-mail: ain.raal@ut.ee
a
Department of Food Science and Hygiene, Estonian University of Life Sci-
ences, Kreutzwaldi 58A, 51014 Tartu, Estonia
b
Institute of Pharmacy, University of Tartu, Nooruse 1, 50411 Tartu, Estonia