Introduction
In adults, detection of any change in acoustic
surroundings activates auditory cortices in temporal
lobes and in frontal brain areas at around 100 ms and
evokes an event-related potential (ERP) called N100.
1
Several neural processes elicit overlapping ERP
components when the sound is novel,
2
when the
sound is attended,
3
and even when there is a deviance
in auditory input which is not consciously observed
(mismatch negativity at 100–200 ms; MMN).
4
Any
novel sensory input also elicits a non-specific vertex
response in the same latency range. This cerebral
response belongs to orienting reaction (OR), which
redirects attention involuntarily to the new stimulus
5
and may set up a neuronal model of stimulus prop-
erties with its first representation.
1,6
Thus, the devel-
opment of a neuronal representation upon stimulus
repetition should inhibit OR and abate corresponding
ERP components. In addition, rapid repetition of
identical stimuli diminishes the size of auditory
ERPs, especially N100, due to refractory properties
of eliciting neuronal populations.
7,8
In children, shortening of ERP latencies with age
is usually taken as a sign of increased efficiency in
receiving networks,
9
while changes in topographic
distribution may indicate developmental changes in
the mode of input processing.
10,11
Auditory ERPs,
which are considered to represent similar cerebral
activation as an ‘adult’ N100, emerge in early infancy
at around 200 ms and the latency shortens until
early adulthood. In a recent study of infants, spoken
syllables evoked first ERP components over temporal
brain areas at 200 ms and these decreased in size
when the stimulus was repeated.
12
Responses at
400 ms diminished similarly but recovered
completely with a change in stimulus. It was
suggested that acoustic analysis was at the level of
detection at 200 ms, whereas auditory features were
recognized in less than 400 ms.
We compared ERPs to elementary auditory input
in school-aged children and in young adults during
infrequent presentation of trains of identical tones.
These trains of tones were separated by silent periods
exceeding the putative duration of echoic mem-
ory.
13,14
The aim of the study was to evaluate auto-
matic processing of ‘novel’ auditory stimuli (i.e. the
first tone in train) and refractory properties of
cerebral responses during the very first repetitions.
Since ERPs to frequent auditory stimulation change
considerably during development, we expected to
observe differential changes in the refractory behav-
iour of ERP components due to different maturation
rates of underlying neuronal populations.
Materials and Methods
We studied 42 healthy school-aged children (mean
age 9 years ± 2 weeks) and 16 healthy young adults
(mean age 28 years, range 23–49 years). All subjects
gave their informed consent before the study.
Tone pips (frequency 800 Hz; duration 85 ms)
were delivered to the right ear in trains of four with
an interstimulus interval (ISI) of 1 s and intertrain
interval (ITI) of 12 s. Identical tones were delivered
with a standard ISI of 1 s during MMN paradigm, in
Neurophysiology, Basic and Clinical
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© Rapid Science Publishers Vol 8 No 6 14 April 1997 1327
WE compared event-related responses (ERPs) to non-
attended frequent and intermittent auditory input in
school-aged children and in young adults. In adults, both
inputs elicited prominent auditory N100 responses at
vertex. In children, intermittent stimulation evoked
vertex responses with similar latency and refractoriness,
whereas frequently delivered identical tones evoked
responses on average at 240 ms. Sensitization of a sepa-
rate neuronal population at 260–300 ms was obvious
during intermittent stimulation in children. The dual
behaviour, simultaneous ‘habituation’ of one neuronal
population response and sensitization of another, may
reflect the process of redirecting the attention and
setting up a neuronal model. Furthermore, results sug-
gest that a simplistic interpretation of developmental
ERPs in which shortening of latencies represents matu-
ration is insufficient.
Key words: Auditory N100; Children; Development;
Event-related potentials; Habituation
Dual cerebral processing
of elementary auditory
input in children
Jari Karhu,
CA
Eila Herrgård,
1
Ari Pääkkönen, Laila Luoma,
1
Eila Airaksinen
1
and Juhani Partanen
Department of Clinical Neurophysiology and
1
Department Pediatrics, Division of Children’s
Neurology, Kuopio University Hospital, Kuopio,
Finland
CA
Corresponding Author
NeuroReport 8, 1327–1330 (1997)