Healthy Aging and Latent Infection with CMV Lead to Distinct Changes in CD8 and CD4 T-Cell Subsets in the Elderly Birgit Weinberger, Lutfan Lazuardi, Ilka Weiskirchner, Michael Keller, Christoph Neuner, Karl-Heinz Fischer, Ber Neuman, Reinhard Würzner, and Beatrix Grubeck-Loebenstein ABSTRACT: Despite general acceptance that immuno- logic changes are associated with aging and latent in- fection with Cytomegalovirus (CMV), no clear-cut distinction has so far been made between strictly age- related and CMV-induced changes. We therefore com- pared CD4 + and CD8 + naïve (CD45RA+CD28+), memory (CD45RA-CD28+), and effector (CD28 - )T cells in CMV-positive (n = 164) and CMV-negative (n = 87) elderly persons and correlated CD8 + and CD4 + effector T cells with other T-cell subpopulations. Per- centages of CD8 + as well as CD4 + effector T cells were higher, but percentages of naïve and memory cells were lower in CMV-positive compared to CMV-negative el- derly persons. Negative correlations within CD8 + T- cell subsets were found to be present in both CMV- positive and CMV-negative elderly individuals. In contrast, correlations within CD4 + T-cell subpopulations and a positive correlation between CD8 + and CD4 + effec- tor T cells were found in CMV-positive individuals only. Our results demonstrate that (a) in the elderly different T-cell subsets compete for space within the CD8 + , but not the CD4 + T-cell population; (b) CMV induces changes in the CD4 + compartment that differ from the solely age- related changes seen in CMV-negative elderly population; and (c) the CMV-status of a population has to be taken into account before a conclusion on the effect of aging on the composition of the T-cell pool can be reached. Human Immunology 68, 86 –90 (2007). © American Society for Histocompatibility and Immunogenetics, 2007. Published by Elsevier Inc. KEYWORDS: CMV; T-cell subsets; aging; CD8 + ; CD4 + ABBREVIATIONS CMV Cytomegalovirus CD cluster of differentiation FACS fluorescence-activated cell sorting IFNInterferon INTRODUCTION It is generally recognized that the function of the im- mune system declines with age [1, 2] leading to in- creased susceptibility to infectious disease and endanger- ing the protective effect of vaccination [3, 4]. Because of the involution of the thymus, changes in the composition of the T-cell repertoire are among the most striking features of immunosenescence. A decreased thymic out- put of naïve T cells leads to low naïve T-cell numbers in the peripheral blood as well as in the lymph node tissue, whereas there is an increase in the numbers of memory and effector T cells [5, 6]. These changes seem to be more pronounced in the CD8 + than in the CD4 + T-cell pool. Longitudinal studies suggest that decreased CD4 + /CD8 + ratios, together with increased numbers of CD57 + and CD28 - T cells and poor T-cell proliferation are part of the “immunologic risk phenotype” (IRP), which corre- lates with subsequent 2-year mortality in the elderly [7, 8]. The pronounced accumulation of CD28 - CD57 - effector cells that produce large amounts of IFNmay From the Immunology Division (B.W., L.L., I.W., M.K., B.G.-L.), Institute for Biomedical Aging Research, Austrian Academy of Sciences, Innsbruck, Austria, Department of Public Health (C.N. K.-H. F.), County of Tyrol, Innsbruck, Austria, Department of Public Health (B.N.), Town- ship of Innsbruck, Innsbruck, Austria, and Department of Hygiene (R.W.), Microbiology and Social Medicine, Medical University, Innsbruck, Austria. Address reprint requests to: Prof. Beatrix Grubeck-Loebenstein, Institute for Biomedical Aging Research, Rennweg 10, 6020 Innsbruck, Austria; Tel: +43-512-583919-10; Fax: +43-512-583919-8; Email: beatrix. grubeck@oeaw.ac.at. B.W. and L.L contributed equally to this paper. Received August 18, 2006; revised October 16, 2006; accepted October 31, 2006. Human Immunology 68, 86 –90 (2007) © American Society for Histocompatibility and Immunogenetics, 2007 0198-8859/07/$–see front matter Published by Elsevier Inc. doi:10.1016/j.humimm.2006.10.019