978-1-4673-1572-2/12/$31.00 ©2012 IEEE
Abstract — Influence of small time-delays in coupling
between noisy excitable systems on the coherence resonance and
self-induced stochastic resonance is studied. Parameters of
delayed coupled deterministic excitable units are chosen such that
the system has only one attractor, namely the stationary state, for
any value of the coupling and the time-lag. Addition of white
noise induces qualitatively different types of coherent oscillations,
and we analyzed the influence of coupling time-delay on the
properties of these coherent oscillations. The main conclusion is
that time-lag 1, but still smaller than the refractory period,
and sufficiently strong coupling drastically change signal-to-noise
ratio in the quantitative and qualitative way. An interval of noise
values implies quite large signal to noise ratio and different types
of noise induced coherence are greatly enhanced. We also
observed coincident spiking for small noise intensity and time-lag
proportional to the inter-spike interval of the coherent spike
trains. On the other hand, time-lags 1 and/or weak coupling
induce negligible changes in the properties of the stochastic
coherence.
— Neurons, Delay, Noise
I. INTRODUCTION
xcitability is a common property of many physical and
biological systems. Although there is no unique definition
[1] the intuitive meaning is clear: a small perturbation
single stable stationary state can result in a large and long
lasting excursion away from the stationary state before the
system is returned back asymptotically to equilibrium. Typical
example of excitable behavior is provided by the dynamics of
neurons. However, realistic models of coupled neurons must
include the following two phenomena: (a) influence of
different types of noise and (b) different time scales of the
creation of impulses on one hand and their transmission
between neurons on the other. It is well known that neurons in
vivo are function under influences of many sources of noise
N. lgrade
68, 11080 Beograd-Zemun, Serbia, e-mail: buric@ipb.ac.rs.
I. and N.Vasovi
Mathematics, Faculty of Mining and Geology, University of Belgra
Box 162, Belgrade, Serbia.
K.
.
is with the Faculty of Transport and Traffic Engineering,
Universi 30 .
[2]. It is also well known that the noise of an appropriate small
intensity can change the systems dynamics by turning the
quiescent state of the neuron into the state of periodic firing
[3]. There are different types of noise induced coherent
oscillations [ , ] that could occur in examples of excitable
systems without [6] or with time-delay [7, 8], as will be
discussed later. Description of interactions between neurons
should include the details of the electrochemical processes in
real synapses which occur on much slower time scale then the
occurrence of an impulse and its transport along axons [9].
Alternatively, the transport of information between neurons
can be phenomenologically described by time-delayed inter-
neuronal interaction. It is well known that, depending on the
parameters, the time-delay can, but need not, induce drastic
qualitative changes on the evolution of coupled deterministic
excitable systems (please see for example [10– ]). However,
a system of delay-differential equations is infinite dimensional
with initial states represented by a vector functions on the
interval ( 0). Stability of stochastic delay-differential
equations has been studied by mathematicians [16, 17].
Influence of noise on time-delay induced bifurcations and
properties of synchronization have been analyzed elsewhere,
for example in [18–22]. On the other hand the influence of
coupling delay on different types of coherent oscillations that
have been induced solely by the noise has been studied much
less [23–
different attractors of systems with time-delayed feedback has
been studied in [26, 27]. Such an analyzes would supply
information complementary to the research on the effects of
noise on the properties of oscillations and synchrony
introduced by sufficient time-lag in the delayed coupling. It is
our goal in this paper to study the effects of time-delay in the
coupling between two excitable units on different types of
noise induced coherent oscillations in each of the units.
II. THE MODEL
In this paper we shall consider typical type II excitable
systems, as modeled by the FitzHugh-Nagumo (FHN)
differential equation [1], where the excitable behavior
bifurcates into the oscillatory regime via the Hopf bifurcation.
white noise that could appear in the model equations in two
qualitatively different ways. Thus each neuron is described by
the following stochastic differential equations:
Coherent Oscillations in Minimal Neural
Network of Excitable Systems Induced by
Noise and Influenced by Time Delay
Nikola Buri
E