Euphytica 127: 335–344, 2002.
© 2002 Kluwer Academic Publishers. Printed in the Netherlands.
335
Obtention of haploid embryos and plants through irradiated pollen
technique in squash (Cucurbita pepo L.)
E.S. Kurtar
1
, N. Sarı
2
& K. Abak
2
1
Ondokuz Mayıs University, Professional High School of Bafra, 55410, Samsun, Turkey;
2
Çukurova University,
Faculty of Agriculture, Department of Horticulture, 01330, Adana, Turkey
Received 28 December 2000; accepted 12 February 2002
Key words:
60
Co, haploidy, irradiated pollen, ploidy determination, squash
Summary
The influence of gamma ray doses (25, 50, 75, 100, 200, 300 and 400 Gy) and genotypes (Eskenderany F
1
, Acceste
F
1
, Sakız, Urfa Yerli) on the induction of haploid embryos obtained by irradiated pollen technique was studied in
squash (Cucurbita pepo L.). Different shapes and stages of embryos were derived from seeds extracted from fruits
harvested 4–5 weeks after pollination. As a result of the present study, haploid embryos and haploid plants were
obtained, with haploid production strongly influenced by gamma ray doses, embryo stages and genotypes. Gamma
ray doses of 25 and 50 Gy gave the highest parthenogenetical response. All of the point shape, globular shape, arrow
tips and stick-shaped embryos developed into haploid plants. However, only 53.8% of torpedo and 23.1% of heart-
shaped embryos gave haploid plants. In contrast, cotyledon-shaped and amorphous-shaped embryos produced only
diploid plantlets. The number of embryos per 100 seeds was the highest in ‘Eskenderany F
1
’ and ‘Sakız’ genotypes.
After in vitro culture, a total of 93 haploid plantlets were obtained.
Introduction
Haploidization techniques facilitate the production of
pure lines from heterozygous plants in a single gener-
ation and represent significant advantages for breeders
and geneticists. Haploid plants can also be obtain
spontaneously, but the frequency of this event is
very low (Pochard & Dumas de Vaulx, 1971). At
the present, haploid plants are produced by anther-
microspore culture and ovule-ovary culture, or by
pollination with incomplete pollen (treated with chem-
icals, irradiated etc.).
Spontaneous haploidy occurs rarely in squash (Cu-
curbita pepo L.) as in other species of the Cucur-
bitaceae family (Savin et al., 1988). Attempts to
produce haploids by in vitro androgenesis and gyno-
genesis in squash were unsuccessful (Chambonnet and
Dumas de Vaulx, 1985; Shail & Robinson, 1987). Re-
cently, gynogenic haploid embryo induction and plant
obtention has been successful through irradiated pol-
len technique in melon (Sauton & Dumas de Vaulx,
1987; Sarı et al., 1992; Maestro-Tejada, 1992; Cuny,
1992; Abak et al., 1996), cucumber (Truong-Andre,
1988; Sauton, 1989; Niemirowicz-Szczytt & Dumas
de Vaulx, 1989; Ça˘ glar and Abak, 1999) and water-
melon (Gürsöz et al., 1991; Sarı et al., 1994). Pollen
irradiation was proved to be effective for haploid in-
duction in melon, but the embryo yield was found to
be highly influenced by different factors such as gen-
otype, environmental conditions and irradiation doses
(Ficcadenti et al., 1995).
The aim of this research was to investigate the
possibility of inducing haploid embryos of Cucurbita
pepo L. through irradiated pollen technique.
Material and methods
Plant material
As plant material, two F
1
hybrids (Eskenderany F
1
and Acceste F
1
) and two local open-pollinated gen-
otypes (Sakız and Urfa Yerli) were used. The seeds
were sown with 30-day intervals, starting from the