VOL. 2, NO. 3, April 2012 ISSN 2225-7217
ARPN Journal of Science and Technology
©2011-2012. All rights reserved.
http://www.ejournalofscience.org
166
Corticosterone as Inhibitor of
Bufo Melanostictus Tadpole Growth
Ruqaiya Hasan and Mahmood Ahmad
Department of Physiology, University of Karachi, Karachi – 75270, Pakistan
Email: ruqaiya55@gmail.com
ABSTRACT
This investigation reports the effect of corticosterone on the growth of Bufo melanostictus tadpoles. Test tadpoles of
experiment I (T-I), maintained in 1μg/ml solution of corticosterone, were 2.39mm shorter than controls by day 11. Test
tadpoles of experiment II (T-II), maintained in 5μg/ml solution were 4.32mm shorter than their counterparts. Growth
inhibition was dose related and became significant (p<0.05) when the larger amount was administered.
Keywords: Metamorphosis, Macrophages, Proteolysis, Anuran tadpoles, Growth inhibitor.
1. INTRODUCTION
Only a minute amount of corticosterone has
been detected [1] in the blood of Amphiuma. Equally, a
very small quantity has been found in the blood of the
adult American bullfrog, Rana catesbeiana [2]. Despite
these demonstrations, it has not yet been shown that
corticosterone is present in the blood of amphibian
larvae [3]. However, several studies have demonstrated
the influence of exogenous corticosterone on the growth
and development of larval amphibians [4, 5, 6].
Kobayashi [4] studied the effects of
deoxycorticosterone acetate on metamorphosis of
thyroxine induced tadpoles. He also claimed that
corticosteroids synergise the destructive phase of
shrinking of tails and a shortening of structures.
The purpose of the present investigation is to
investigate the destructive action of corticosterone in
premetamorphic Bufo melanostictus tadpoles.
2. MATERIALS AND METHODS
Animals:
Bufo melanostictus tadpoles belonging to a
single hatching were collected from the rainwater ponds
of the University Campus in the month of August 2011.
After collection, tadpoles of similar size were divided
into several groups of equal number. They were soon
given refined suspension of parboiled spinach [7, 8].
They were then allowed to acclimatize for a number of
days. On attaining premetamorphic period, tadpoles
were used to determine the effect of corticosterone; since
physiological connections between pituitary and
hypothalamus are established when they enter
premetamorphic period of stage 52 [9, 10, 11]. The
tadpoles were selected with the help of a binocular
microscope; and after measurement, individuals of the
same size were randomly divided into comparable
groups of control and test.
Growth correlations:
It may be stated that during development, there
appears to be gradual unfolding with time; of anterior
pituitary and receptor-gland hormones that regulate the
final size, form and body parts. This elaboration of
growth is indicative of a second phase of specific
constitutional growth potential, provided by the
genotype of the egg. During this phase the growth
regulating hormones gradually take over the function of
the coordination and regulation of final size [12].
However, growth is also dependent upon environmental
factors [13]; since nutrients, oxygen supply [14] and
temperature have an effect on the physico-chemical
background in which the synthetic processes of growth
take place. Thus, growth follows the expression of size
and weight. Therefore, growth of Bufo melanostictus
tadpoles was worked out by measuring its body size and
decisive evidence of growth has also been sought
through the use of these measurements [15].
Method of measurement:
The method adopted for this purpose was
similar to that described by Ahmad and Mukarram [16];
and was later adapted by Ahmad et al., [15, 17, 18, 19].
In order to measure each tadpole, nylon net was used to
introduce tadpoles into a 6 cm diameter glass Petri dish,
in the absence of water. As soon as the tadpole ceased its
wriggling movements, the Petri dish was tilted slightly
towards the head side. By sliding, the tadpole body was
shaped into a straight line. The Petri dish was arranged
on a glass bearing a millimeter grid and the length from
the head to tail was noted.
Design of experiment:
Space is a major factor in the growth rate of
tadpoles, and crowding has been reported to accelerate
metamorphosis by operating through the hypothalamic