© 2008 Wiley-VCH Verlag GmbH & Co. KGaA, Weinheim 871
Biotechnol. J. 2008, 3, 871–877 DOI 10.1002/biot.200700197 www.biotechnology-journal.com
1 Introduction
In the last several years, fundamental studies have
examined a new field of research concerned with
the exploitation of novel environmental and eco-
logical materials derived from marine wastes. Im-
mense quantities of marine processing byproducts
including fish bone, fish muscle protein and inter-
nal organs, fish skin collagen and gelatin, fish oils
such as omega-3 fatty acids, shellfish and crus-
tacean shell wastes are of both environmental and
economic concern for seafood processing plants
[1]. At present, current discards from the world’s
fisheries exceed 20 million tons, which equates to
almost 25% of the total production of marine cap-
ture fisheries [2]. Moreover, European legislation
(EU Council Directive 1999/31/EC, 1999) has set
specific targets for the amount of biodegradable
municipal waste (BMW) for disposal at landfills [3].
In Ireland, figures from An Bord Iascaigh Mhara
(The Irish Sea Fisheries Board), estimate that
63,786 tonnes of marine processing waste were
produced nationally in 2002, 45% of which arose
from the northwest region [4]. Traditionally, the
greater part of fisheries wastes are utilized in the
production of low-economic value products such as
fertilizers, fish silage or pet foods [5]. However, with
advances in bioprocess engineering technologies
coupled with novel enzymatic and microbial hy-
drolysis methods, processing wastes may serve as
cheap raw materials for the generation of high-val-
ue bioactive compounds with a variety of applica-
tions [1, 6, and 7]. Various applications of chitin,
Review
Mining marine shellfish wastes for bioactive molecules:
Chitin and chitosan – Part A: extraction methods
Maria Hayes
1
, Brian Carney
1
, John Slater
1
and Wolfram Brück
1
1
Centre of Applied Marine Biotechnology (CAMBio), Letterkenny Institute of Technology, Port Road, Letterkenny, Co. Donegal,
Ireland.
Legal restrictions, high costs and environmental problems regarding the disposal of marine pro-
cessing wastes have led to amplified interest in biotechnology research concerning the identifica-
tion and extraction of additional high grade, low-volume by-products produced from shellfish
waste treatments. Shellfish waste consisting of crustacean exoskeletons is currently the main
source of biomass for chitin production. Chitin is a polysaccharide composed of N-acetyl-D-glu-
cosamine units and the multidimensional utilization of chitin derivatives including chitosan, a
deacetylated derivative of chitin, is due to a number of characteristics including: their polyelec-
trolyte and cationic nature, the presence of reactive groups, high adsorption capacities, bacterio-
static and fungistatic influences, making them very versatile biomolecules. Part A of this review
aims to consolidate useful information concerning the methods used to extract and characterize
chitin, chitosan and glucosamine obtained through industrial, microbial and enzymatic hydrolysis
of shellfish waste.
Keywords: Chitin · chitosan · oligosaccharides · bioactive molecules
Correspondence: Dr. Brian Carney, Centre of Applied Marine Biotechnolo-
gy CAMBio, Letterkenny Institute of Technology, Port Road, Letterkenny,
Co. Donegal, Ireland.
E-mail: Brian.Carney@Lyit.ie
Telephone: +353 (0) 74 9186565
Fax: +353-749186300
Abbreviations: COS, chitosan oligosaccharides, DCE, cis-1,2-dichloro-
ethene, DD, degree of deacetylation, DP, degrees of polymerisation, ECM,
extracellular matrix, FITC-chitin, Fluorescein isothiocyanate chitin, GlcNAc,
N-acetylglucosamine, LMMC, low molecular mass chitosans, NOCC, N-O-
Carboxymethyl chitosan, PVC, polyvinyl chloride, SCSP, shrimp and crab
Received 22 August 2007
Revised 24 January 2008
Accepted 31 January 2008