Pharmacology Biochemistry & Behavior, Vol. 9, pp. 735-740. Printed in the U.S.A.
Cycloheximide Produces Attentional
Persistence and Slowed Learning in Chickens
L.J. ROGERS
Pharmacology Department, Monash University, Clayton, Victoria, 3168, Australia
AND
J. M. ANSON
Department of Behavioural Biology, Australian National University, Canberra, Australia
(Received 17 February 1978)
ROGERS, L. J. AND J. M. ANSON. Cycloheximide produces attentional persistence and slowed learning in chickens.
PHARMAC. BIOCHEM. BEHAV. 9(6) 735-740, 1978.--Cycloheximide has been used previously to demonstrate that
long-term memory has a protein substrate. We have shown that it has two additional behavioural effects when it is
administered to young chickens. A dose of 10/zg of cycloheximide in 5 p.l of normal saline vehicle given to each side of the
forebrain on Day 2 of post-hatched life changes attention so that responding to a given stimulus type becomes more
persistent. Behavioural testing occurred in the chickens' second week of life. They were given two colours of food, and
runs of pecks at each colour were scored. Treated chicks showed less frequent switching from one colour to the other. A 20
/~g dose of cycloheximide in 25/~1 of saline also produced this state of attentional persistence, and additionally it made the
chickens slower at learning a visual discrimination task, a visual habituation task and an auditory habituation task. The
paper concludes that the effect on attention is separate from that on learning, and discusses the findings with reference to
testosterone's known ability to produce attentional persistence and also with reference to previous research with cyc-
loheximide and long-term memory.
Cycloheximide Attentional persistence
Visual discrimination learning Chickens
Slowed learning Long-term memory Retrieval
CYCLOHEXIMIDE, a drug which blocks ribosomal protein
synthesis, has been used frequently in studies on memory
formation, where it is claimed to block consolidation of long
term memory [5, 6, 8, 13]. In addition to this property, cy-
cloheximide has been found to produce a permanent reduc-
tion in learning rate when it is administered to young chick-
ens [12]. When administered intracranially to a chicken in
the first week of life, it not only blocks the formation of long
term memory of tasks performed about the time of treat-
ment, but it also alters the brain in such a way that learning
on a number of visual and auditory tasks is found to be
permanently retarded.
Our previous studies have almost always used a 20/~g
dose of cycloheximide dissolved in 25/~1 of normal saline
administered to each side of the forebrain [12,13]. We there-
fore considered it important to measure the amount of learn-
ing retardation produced by varying the amounts of cy-
cloheximide administered in various volumes of saline, and
from these studies an interesting new effect of cycloheximide
on behaviour emerged.
Cycloheximide was found to cause the chicken to persist
in responding to a given stimulus type once it had begun
responding to that stimulus type. This appeared to be re-
markably similar to the persistence in responding, or atten-
tional persistence, found previously in male chickens treated
with testosterone [4,11]. It was therefore studied in visual
search tests designed to measure attentional persistence by
scoring runs of pecking on differently coloured food. By
appropriate choice of dose followed by measuring both learn-
ing rate on a visual discrimination task and attentional persis-
tence, it was possible to separate learning effects from
those of attentional persistence.
LEARNING RATES
METHOD
Housing
White leghorn-black australorp cross male chickens were
obtained from a commercial hatchery on Day 1 of life. They
were housed in groups of 4-6 birds until the evening of Day 3
when they were separated visually from each other. The
cages (23x23 cm sq. and 29 cm high) were made of metal
with a clear perspex front, through which the chickens could
see the daily activity in the laboratory and become
familiarized with the experimenter. Constant light and
warmth were provided by a 25 W globe suspended above the
cage. (For further details see [14]).
Feeding
Food and water were freely available and changed reg-
Copyright © 1978 ANKHO International Inc.--0091-3057/78/120735-06501.10/0