EXPERIMENTAL STUDIES
A High Resolution Assessment of the Repeatability of Relative
Location and Intensity of Transcranial Magnetic
Stimulation–induced and Volitionally Induced Blood Oxygen
Level–dependent Response in the Motor Cortex
Stewart Denslow, PhD,* Mikhail Lomarev, MD, PhD,*¶ Daryl E. Bohning, PhD,*
Qiwen Mu, MD, PhD,* and Mark S. George, MD*†‡§
Objective: Using functional magnetic resonance imaging, we as-
sessed variation in location and intensity of blood oxygen level–
dependent contrast associated with movements induced by transcra-
nial magnetic stimulation or volition.
Background: Anatomic location and within-subject repeatability
of blood oxygen level–dependent responses induced by transcranial
magnetic stimulation comprise critical information to the use of in-
terleaved transcranial magnetic stimulation/functional magnetic reso-
nance imaging as a neuroscience tool.
Methods: Eleven healthy adults were scanned 3 times each at 1.5 T.
Interleaved with functional magnetic resonance imaging, 1-Hz trans-
cranial magnetic stimulation was applied over motor cortex. VOL
was alternated with transcranial magnetic stimulation over the scans.
Results: Intra-subject standard deviations in blood oxygen level–
dependent locations ranged between 3 and 6 millimeters, allowing
localization to subregions of the motor strip. Coil placement relative
to blood oxygen level–dependent location varied more than blood
oxygen level–dependent location (sd
x
= 9.5mm, sd
y
= 8.7mm, sd
z
=
9.0mm) with consistent anterior displacement (dy = 21.8 mm, P =
<0.025). Analysis of variance did not detect significant differences
between transcranial magnetic stimulation and VOL blood oxygen
level–dependent locations or intensities, in contrast to significant inten-
sity differences detected in auditory blood oxygen level dependence.
Conclusion: The high repeatability of location of transcranial mag-
netic stimulation–induced blood oxygen level–dependent activation
suggests that transcranial magnetic stimulation/functional magnetic
resonance imaging stimulation can be used as a precise tool in inves-
tigation of cortical mechanisms. The similarity between VOL and
transcranial magnetic stimulation suggests that transcranial magnetic
stimulation may act through natural brain movement circuits.
Key Words: functional magnetic resonance imaging, human motor
cortex, volitional movement, blood flow
(Cog Behav Neurol 2004;17:163–173)
T
he capacity of transcranial magnetic stimulation (TMS) to
directly stimulate cortical neurons
1
has made TMS attrac-
tive as a probe of basic neural interconnections.
2,3
The nonin-
vasive nature of TMS has also generated great interest in its use
for investigation and treatment of depression.
2,4,5
These fac-
tors have led to growing interest in combining TMS with func-
tional imaging methods to explore brain circuitry.
6–12
Both
positron emission tomography and functional magnetic reso-
nance imaging (fMRI) observations have shown similarity in
the brain’s responses to TMS and cognitive tasks.
6,7,9,10,13
However, the relations among cortical response locations,
TMS coil locations, and specific cortical anatomy are not yet
precisely described. Additionally, recent fMRI results have
demonstrated the large variability in volume of blood oxygen
level dependent (BOLD) activation that is encountered be-
tween repeated scans.
14,15
It is consequently unclear what
magnitude of variation in location indices can be expected for
BOLD responses both from physiological tasks and from TMS
stimulation.
TMS depolarization of axons displays a complex depen-
dence upon configuration between the induced electric field
and axonal axis. Models have shown greater likelihood of de-
polarization for axons parallel rather than perpendicular to the
Received for publication June 2, 2003; revised October 10, 2003; accepted
November 16, 2003.
From the *Center for Advanced Imaging Research and Brain Stimulation
Laboratories, Department of Radiology, Medical University of South
Carolina, Charleston, South Carolina; †Center for Advanced Imaging Re-
search and Brain Stimulation Laboratories, Department of Psychiatry,
Medical University of South Carolina, Charleston, South Carolina; ‡Cen-
ter for Advanced Imaging Research and Brain Stimulation Laboratories,
Department of Neurology, Medical University of South Carolina, Charles-
ton, South Carolina; §Ralph H. Johnson Veterans Hospital, Charleston,
South Carolina; and ¶Institute of the Human Brain, St. Petersburg, Russia.
Supported in part by research grants from the Dana Foundation (Bohning),
from the National Institute of Neurological Disorders and Stroke (RO1
RR14080-02), and from the Defense Advanced Research Projects Agency
Defense Sciences Office. None of the authors have equity or significant
financial conflicts.
Reprints: Dr. Stewart Denslow, Radiology Department, 171 Ashley Avenue,
Medical University of South Carolina, Charleston, SC 29425 (e-mail:
denslows@musc.edu).
Copyright © 2004 by Lippincott Williams & Wilkins
Cog Behav Neurol • Volume 17, Number 3, September 2004 163