Fungal Morphology and Fragmentation
Behavior in a Fed-Batch Aspergillus
oryzae Fermentation at the
Production Scale
Zheng Jian Li,
1
Vivek Shukla,
1
Andrew P. Fordyce,
2
Annemarie Gade Pedersen,
2
Kevin S. Wenger,
3
Mark R. Marten
1
1
University of Maryland, Baltimore County (UMBC), Department of
Chemical and Biochemical Engineering, 1000 Hilltop Circle, Baltimore,
Maryland 21250; telephone (410) 455-3439; fax (410) 455-1049; E-mail:
marten@umbc.edu
2
Novo Nordisk A/S, Enzyme Research, Bagsvaerd, Denmark
3
Novo Nordisk BioChem North America, Inc., Franklinton, North Carolina
Received 22 November 1999; accepted 14 May 2000
Abstract: It is well known that high-viscosity fermenta-
tion broth can lead to mixing and oxygen mass transfer
limitations. The seemingly obvious solution for this
problem is to increase agitation intensity. In some pro-
cesses, this has been shown to damage mycelia, affect
morphology, and decrease product expression. How-
ever, in other processes increased agitation shows no
effect on productivity. While a number of studies discuss
morphology and fragmentation at the laboratory and pi-
lot scale, there are relatively few publications available
for production-scale fungal fermentations. The goal of
this study was to assess morphology and fragmentation
behavior in large-scale, fed-batch, fungal fermentations
used for the production of protein. To accomplish this, a
recombinant strain of Aspergillus oryzae was grown in
80 m
3
fermentors at two different gassed, impeller
power-levels (one 50% greater than the other). Impeller
power is reported as energy dissipation/circulation func-
tion (EDCF) and was found to have average values of 29.3
± 1.0 and 22.0 ± 0.3 kW m
–3
s
-1
at high and low power
levels, respectively. In all batches, biomass concentra-
tion profiles were similar and specific growth rate was
< 0.03 h
-1
. Morphological data show hyphal fragmenta-
tion occurred by both shaving-off of external clump hy-
phae and breakage of free hyphae. The fragmentation
rate constant (k
frag
), determined using a first-order
model, was 5.90 and 5.80 h
-1
for high and low power
batches, respectively. At the end of each batch, clumps
accounted for only 25% of fungal biomass, most of which
existed as small, sparsely branched, free hyphal ele-
ments. In all batches, fragmentation was found to domi-
nate fungal growth and branching. We speculate that this
behavior was due to slow growth of the culture during
this fed-batch process. © 2000 John Wiley & Sons, Inc. Bio-
technol Bioeng 70: 300–312, 2000.
Keywords: filamentous fungi; fermentation, morphol-
ogy; fragmentation; agitation
INTRODUCTION
Filamentous fungi offer many benefits over other types of
cells when used for the production of chemicals and phar-
maceuticals and, as a result, are used to produce products
comprising a large fraction of the world’s pharmaceutical
and biotechnology market (Arora et al., 1992). The wide-
spread use of fungi in biotechnology stems from the unique
ability of fungi to economically produce many different
types of products, including some of the most important
antibiotics (e.g., penicillins and cephalosporins; Masurekar,
1991; Rambosek, 1991), commodity chemicals (e.g., citric
acid; Zidwick, 1991), and commercial enzymes (e.g., pro-
teases and amylases; Bigelis, 1991; Arora et al., 1992). For
the production of enzymes and therapeutic proteins, fungi
offer several advantages over other types of cells. First
among these is the large amount of protein (greater than 50
g/L in some cases) fungi are able to both produce and se-
crete to the growth medium. In addition, fungi generally
exhibit correct glycosylation/posttransitional modification
of proteins, are often classified as GRAS (generally re-
garded as safe) by the FDA (Food and Drug Administra-
tion), and can be grown on a wide variety of inexpensive
substrates. In addition, recent improvements in host cells
and expression vectors enable filamentous fungi to be con-
sidered as a practical alternative to bacteria, yeast, insect, or
mammalian expression systems for production of heterolo-
gous gene products (Fowler and Berka, 1991).
While filamentous fungi have been used for many years
to produce a vast array of products, many fungal fermenta-
tions suffer from the same problem—high broth viscosity
that often leads to oxygen mass-transfer limitations (Morris
et al., 1973; Metz et al., 1979; Atkinson and Mavituna,
1991; McNeil and Harvey, 1993; Olsvik and Kristiansen,
1994). In a mechanically agitated tank, one frequently used
method to overcome oxygen limitations is simply to in-
crease impeller power, thus increasing oxygen mass-
Correspondence to: M.R. Marten
Contract grant sponsor: the National Science Foundation
Contract grant number: BES-9876012
© 2000 John Wiley & Sons, Inc.