Journal of Reproductive Immunology 53 (2002) 161–169 Signaling through protein kinases during egg activation Efrat Eliyahu, Anat Talmor-Cohen, Ruth Shalgi * Department of Embryology, Sackler Faculty of Medicine, Tel AiUniersity, Ramat -Ai69978, Tel -Ai, Israel Received 9 April 2001; accepted 16 April 2001 Abstract Following penetration of the zona pellucida, the mammalian spermatozoon binds and fuses with the egg plasma membrane, thereby fertilizing the egg that is still arrested at the second metaphase. Fertilization initiates in the egg a sequence of events referred to as ‘egg activation’. An initial increase in intracellular Ca 2 + concentration ([Ca 2 + ] i ) appears to be the very early cellular event observed which leads to the cortical granules exocytosis and resumption of meiosis. Various treatments can induce parthenogenetic activation mimicking at least part of the fertilization events. Similar to somatic cells, studies in mammalian eggs suggest that signal transduction pathways mediate egg activation. The initial increase in [Ca 2 + ] i appears to be critical for egg activation. However, other messengers such as protein kinase C (PKC) and protein tyrosine kinases (PTKs), were suggested as possible inducers of some aspects of egg activation. In the present work, studies concerning the involvement of protein kinases during egg activation in our laboratory and in others are summarized. © 2002 Elsevier Science Ireland Ltd. All rights reserved. Keywords: Egg; Signal transduction; Activation; Mammals www.elsevier.com/locate/jreprimm 1. Introduction At fertilization, a spermatozoon binds to and then fuses with an egg that is arrested at the second meiotic metaphase. Sperm egg interaction initiates * Corresponding author. Tel.: +972-3-640-8685; fax: +972-3-640-6149. E-mail address: shalgir@post.tau.ac.il (R. Shalgi). 0165-0378/02/$ - see front matter © 2002 Elsevier Science Ireland Ltd. All rights reserved. PII:S0165-0378(01)00097-3