Zoological Journal of the Linnean Society (2000), 128: 77–121. With 12 figures doi: 10.1006/zjls 1998.0184, available online at http://www.idealibrary.com.on The phylogeny of the ‘higher’ temnospondyls (Vertebrata: Choanata) and its implications for the monophyly and origins of the Stereospondyli ADAM M. YATES* AND A. ANNE WARREN School of Zoology, La Trobe University, Bundoora, Melbourne, Victoria 3083, Australia Received March 1998; accepted for publication August 1998 A parsimony analysis of ‘higher’ temnospondyls (all temnospondyls descended from the common ancestor of Eryops and Parotosuchus) was performed using 37 terminal taxa and 121 osteological characters. Bremer support values for each internal node were calculated as a measure of clade strength. Additionally, the shortest trees that conformed to some alternative hypotheses were searched for. The following new taxa are established on the basis of the results: Euskelia (the clade containing the Eryopoidea and Dissorophoidea), Limnarchia (the clade containing Trimerorhachidae, Dvinosauroidea, Archegosauroidea and Stereospondyli), Dvinosauria (the clade containing Trimerorhachidae and Dvinosauroidea), Stereo- spondylomorpha (the clade containing Archegosauroidea and Stereospondyli), Capitosauria (the clade containing Lydekkerina and ‘capitosauroids’), and Trematosauria (the clade containing Trematosauroidea, Rhytidosteidae, Plagiosauroidea, Metoposauroidea and Brachyopoidea). The monophyly of the assemblage of Mesozoic families called the Stereospondyli by Romer is supported. The dominance of the Stereospondyli in the Mesozoic and its rarity in the Palaeozoic is discussed. It is suggested that the radiation of the diverse stereospondyl clades, the Capitosauria and Trematosauria, began in the Late Permian of Gondwana, in a ‘safe haven’ that was less severely affected by the Late Permian extinction event. It is further speculated that the ‘safe haven’ was located in Antarctica, or possibly Australia.  2000 Academic Press ADDITIONAL KEY WORDS:—phylogenetic – Temnospondyli – Gondwana – Permian – Triassic – Palaeozoic – Mesozoic. CONTENTS Introduction . . . . . . . . . . . . . . . . . . . . . . . 78 Definition of the Temnospondyli . . . . . . . . . . . . . . . . 78 Monophyly of the ingroup . . . . . . . . . . . . . . . . . . 79 Material and methods . . . . . . . . . . . . . . . . . . . 80 Abbreviations . . . . . . . . . . . . . . . . . . . . . . 83 Results . . . . . . . . . . . . . . . . . . . . . . . . 84 Discussion . . . . . . . . . . . . . . . . . . . . . . . 104 Acknowledgments . . . . . . . . . . . . . . . . . . . . . 109 References . . . . . . . . . . . . . . . . . . . . . . . 110 Appendix 1 . . . . . . . . . . . . . . . . . . . . . . . 114 Appendix 2 . . . . . . . . . . . . . . . . . . . . . . . 120 * Corresponding author. E-mail: adam.yates@ 77 0024–4082/00/010077+45 $35.00  2000 Academic Press