ELSEVIER Journal of the Autonomic Nervous System 52 (1995) 225-236
Journal of the
Autonomic
Nervous
System
Co-existence of nitrergic, peptidergic and acetylcholine
esterase-positive nerves in the pig lower urinary tract
Katarina Persson a Per Aim b, * Kjell Johansson c, Bengt Larsson a,
Karl-Erik Andersson a
a Department of Clinical Pharmacology, University of Lund, S-221 85 Lund, Sweden
b Department of Pathology, University of Lund, S-221 85 Lund, Sweden
c Department of Zoology, University of Lund, S-221 85 Lund, Sweden
Received 9 February 1994; revision received and accepted 24 August 1994
Abstract
The distribution of NO synthase (NOS) immunoreactive nerves and the possible co-existence with other
neurotransmitters were investigated in the pig lower urinary tract. NOS immunoreactive nerves were found in the
muscle layer, in the lamina propria and around blood vessels. The density of NOS immunoreactive nerves was more
prominent in the trigone and urethra than in the detrusor. All parts of the lower urinary tract were supplied by
numerous acetylcholine esterase (ACHE) positive nerves. The number of adrenergic nerves in the trigone and
urethra was moderate to rich, whereas only very few adrenergic nerves were demonstrated in the detrusor. A low to
moderate number of nerve fibres containing neuropeptide Y (NPY) and vasoactive intestinal polypeptide (VIP) were
observed in the trigone and urethra, while very few were found in the detrusor. A small number of nerves, confined
to the trigone and urethra, were stained for calcitonin-gene-related peptide, somatostatin and leu-enkephalin. Nerve
fibres exhibiting immunoreactivity to bombesin/gastrin releasing peptide, gastrin/cholecystokinin, substance P or
neurokinin A were virtually absent. Co-localization studies revealed that some NOS-immunoreactive nerves also
stained for NPY, VIP or ACHE. The present study shows that nitrergic nerves are present in the pig lower urinary
tract in a density lower than the cholinergic, but higher than any of the studied peptidergic nerves. Coinciding
localization of NOS-positive nerves with nerves expressing ACHE, VIP and NPY suggests that NO may have a role
as a messenger in the lower urinary tract directly and by interaction with other transmitters.
Keywords: Urinary tract; Nitric oxide synthase; Autonomic nervous system; Neuropeptide; (Pig)
I. Introduction
In the urinary tract, much interest has been
focused on peptides as potential mediators of
* Corresponding author. Tel.: (46-46) 173-405; Fax: (46-46)
143-307.
non-adrenergic, non-cholinergic (NANC) effects,
and both intraneuronal storage and functional
responses can be demonstrated (e.g., Refs. 3 and
30). The purine ATP seems to be the main non-
cholinergic excitatory transmitter in the detrusor
[3,10]. Despite extensive research, no candidate
for the inhibitory NANC transmitter involved in
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