Pharmacology B~ochemtstry& Behavior, Vol 26, pp 539-542 Copyright © PergamonJournalsLtd , 1987 Pnntedm the U S A 0091-3057/87 $3 130+ 00
Short-Term and Delayed Behavioral
Effects of Pre- and Post-Weaning
Morphine in Mice I
ENRICO ALLEVA AND GIOVANNI LAVIOLA
Section of Neurobehavloral Pathophyslology, Laboratorlo di Flslopatologia dl Organo e di Sistema
Istituto Superiore di Saint6, V.le Regina Elena, 299, 1-00161 Roma, Italy
Received 15 April 1986
ALLEVA, E AND G LAVIOLA Short-term and delayed behavtoral effects of pre- and post-weamng morphine tn mtce
PHARMACOL BIOCHEM BEHAV 26(3) 539--542, 1987 --Ninety mouse pups of the CD-1 outbred strata were used to
assess actlwty (Vanmex Activity Meter, Columbus Instr, OH) and analgesia (hot plate) after morphine hydrochlonde
given IP either on days 14-16 (preweanhngs) or on days 21-23 (postweanhngs) In preweanhngs morphine depressed
activity already at the lowest dose tested (0 5 mg/kg), and higher doses (1, 5, and 10 mg/kg) d~d not produce a s~gmficantly
larger effect Activity of postweanhngs was not depressed until a very h~gh dose (20 mg/kg) By contrast, morphine
produced clear analgesic effects at all doses m both preweanhngs (day 14) and postweanhngs (day 21) Around day 70,
activity and hot-plate tests m the no-drug state showed no differences due to prior treatment, except for the fact that
hot-plate latenc~es of m~ce previously ~njected wtth sahne as preweanhngs were h~gher than those of all other groups
Twenty-four hr later the tests were repeated after morphine mject~on (10 mg/kg), and showed a s~gmficantly greater
depression of activity m m~ce prewously exposed as preweanhngs On the other hand, all groups previously exposed to
morphine at e~ther the pre- or the post-weanhng stage showed tolerance to the analgesic effect of the drug These
developmental profiles confirm that op~oid systems contribute to the modulation of activity by mechanisms which are at
least m part separate from those mediating analgesia
Mouse behavioral development Morphine analgesia
Environmentallyreduced analgesia (EIA)
Hot plate Locomotor actiwty Adult tolerance
SYSTEMIC admtnistrat~on of morphine to altrlc~al rodents
at d~fferent postnatal stages results in quite different physi-
ological and behavioral changes [6, 14, 15, 17, 19, 20] Speof-
~cally, the analgesic effect of the drug m rats increases from
early postnatal period until about day 15 [6,14] in parallel to a
rapid rise in the number of ~-receptors [19] In the subse-
quent period, drug sens~t~wty declines [6,15] in parallel to the
reduction of general drug toxicity [17], and these changes are
apparently related to the maturation of the blood/brain bar-
her [6,17]
Locomotor actlwty ~s also differentially affected by mor-
phine as a function of postnatal age. One study showed a
reduction of activity ~n rats of ten days, and either no ef-
fects or actiwty enhancements at later ages [9] In another
study, Inbred mice of the C57B1/6 strain showed mmnly
hyperactivity during ontogeny, except for a brief period
(days 20-22) in which the drug produced a marked catatoma
[11]
More recently, attention has been given to the proactive
effects of postnatal morphine exposure on subsequent re-
sponding to the drug For example, extended morphine ad-
ministration ~n rats from birth to weaning produced tolerance
to the analgesic and hypoactivatmg effects of the drug at 22
days w~thout, however, any appreciable effect on
/x-receptors prohferatton [7]. The same investigators also
showed that a single dose of morphine given to newborn rats
sufficed to produce a similar change in sensitivity 26 days
later. Th~s change faded to appear in pups ~njected only a few
days later, that is, e~ther on day 5, 9, or 13 [8].
The present work' was designed to assess the effects of
different doses of morphine on pain sensitivity and locomo-
tor activity of mice before and after weaning, and the conse-
quences of such exposure on adult sensitivity to the same
drug. Animals were treated and tested for three consecutive
days either dunng the last part of the pre-visual stage (pre-
weanhngs, days 14-16, eye opening is completed on day 17
~n the strain used [5]), or starting on the day after weaning
(postweanhngs, days 21-23) These treatment periods were
chosen also on the bas~s of a previous study which showed
that mouse pups of these two ages have quite different pat-
terns of activity, habituation, and response to model drugs
such as amphetamine and scopolamine [2]. All animals were
retested at about 70 days of age for actiwty, pain reactivity,
and morphine effects thereon
~Part of the data on activity and analgesia in ~mmature mice were presented at the IX European Neurosoence Association congress (Oxford,
U K , 8-12 September, 1985) [4]
539