BrainResearchBulletin, Vol. 32, pp. 57-63, 1993 0361-9230/93 $6.00 + .00
Printedin the USA.All rightsreserved. Copyright© 1993Pergamon PressLtd.
Hash and Pattern Reversal Visual Evoked
Potentials in C57BL/6J and B6CBAF /J mice
GEORGE M. STRAIN ~ AND BRUCE L. TEDFORD
Veterinary Physiology, Pharmacology and Toxicology, School of Veterinary Medicine,
Louisiana State University, Baton Rouge, LA 70803-8420
Received 3 August 1992; Accepted 9 February 1993
STRAIN, G. M. AND B. L. TEDFORD. Flashand pattern reversal visualevokedpotentials in C57BL/6J and B6CBAFJJ mice.
BRAIN RES BULL 32(1) 57-63, 1993.--Visual system responses(visual evoked potentials) to flash (FVEP) and pattern reversal
(PRVEP) stimuli were recorded in mice. Two strains were used: black C57BL/6J mice and agouti B6CBAFdJ mice (first generation
offspring of C57BL/6J females and CBA/J males.) Subjects were sedated with ketamine and xylazine. Flash rate (FVEP) and
stimulus spatial frequency and pattern reversal rate (PRVEP) were varied to determine optimum stimulus parameters. Normative
FVEP and PRVEP data were collected from mice of both strains after determination of optimum parameters. Five positive and
four negative alternating peaks were routinely observed in the FVEP, while three positive and three negative alternating peaks
were seen with the PRVEP. Varying the flash rate, the pattern reversal rate, and spatial frequency significantly affected nearly all
amplitude and latency measures in the responses. Significantdifferences between strains were seen on some, but not all, latency
and amplitude measures when the stimulus parameters were varied.
Pattern reversal visual evoked potentials Flash visual evoked potentials B6CBAF~/J vs. C57BL/6J mice
NONINVASIVE objective evaluation of visual pathway function
is most often accomplished through the visual evoked potential
(VEP), where scalp responses are recorded to appropriate visual
stimuli, either flashes of white light or an alternating pattern of
checks or bars. The VEP has been described for numerous spe-
cies, including human (4,6,24), monkey (6,16), dog (26), cat (6),
rat (3,6,19-21), guinea pig (6), cow (27), pig (18), and sheep
(25), among others. Applications of flash VEPs in mice have
been reported in drug (2) and senescence studies (1), but detailed
characterizations of the responses to flash and pattern reversal
stimuli in mice have not been reported. The characterizations
reported here were performed to enable visual system studies of
transgenic mice with disrupted myelinization of the optic nerves.
Two related strains were compared to evaluate within-species
variability. Methods were developed to enable recordings without
implantation of chronic electrodes because of an anticipated
low availability of transgenic subjects.
METHOD
Animals
Young adult male mice (average weight 25 g) from two strains
were used: black mice of the C57BL/6J strain (n = 15) and
agouti mice (B6CBAF~/J, n = 14) which were first generation
offspring of C57BL/6J females and CBA/J males (Jackson Lab-
oratory, Bar Harbor, ME). Retina pigmentation is present in
both mouse strains.
Anesthesia
Mice were lightly anesthetized with a combination of IP ket-
amine and xylazine to avoid the cortical depressant effects of
injectable barbiturate anesthetics; gas anesthetic use was pre-
cluded by the need for unobstructed visual stimulation. The
B6CBAF~/J mice required higher doses (130 and 26 mg/kg, re-
spectively) than the C57BL/6J mice (80 and 16 mg/kg). When
required, supplemental doses of 1/4 the initial dose were ad-
ministered. No mydriatic drugs or dark adaptation were used,
and refraction of the eye was not performed because of the con-
siderable depth of field of rodent eyes (15). During recordings
the mice were placed in a depression on an isothermal chemical
heating pad (Deltaphase, Braintree Scientific, Braintree, MA)
that maintained a constant 37°C temperature to counteract hy-
pothermia effects of the anesthesia.
Visual Stimulation
Recordings were collected in a darkened room with a
background illuminance of 0.25 lumens/ft 2. Stimuli were
binocular, presented in front of the nose, and on a plane level
with the mouse's head. Flash stimuli of 10 us duration from
a photostimulator (Model PS22 photic stimulator, Grass In-
struments Co., Quincy, MA) were presented at a distance of
15 cm from the eyes. A flash intensity setting of 16 was used,
which represented an approximate peak intensity of 2 × 10 7
lumens. Flash rates of 1, 3, 5, and 7/s were evaluated in eight
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