© 2002 Blackwell Science Ltd. http://www.blackwell-science.com/ddi 129
BIODIVERSITY RESEARCH
Diversity and Distributions (2002) 8, 129–145
Blackwell Science Ltd
Distribution of South African marine benthic invertebrates
applied to the selection of priority conservation areas
A. ADNAN AWAD
1
*, CHARLES L. GRIFFITHS
2
and JANE K. TURPIE
1
1
Percy Fitzpatrick Institute, University of Cape Town, Rondebosch, South Africa
2
Department of Zoology,
University of Cape Town, Rondebosch 7701, South Africa
Abstract. Available data on species distributions
and endemicity were compiled and examined for
11 groups of South African marine invertebrates
(2533 species). For five groups species richness
adhered to a well-documented pattern, increasing
from west to east, but for the other groups
species richness was highest along the south
coast. Endemicity was generally highest along
the south coast, and lowest along the east coast.
The data base was then analysed using several
types of complementarity analyses, each produc-
ing a minimum set of potential reserve areas,
which cumulatively represent all invertebrate
species analysed. Approaches based solely on rarity,
species richness and endemicity demonstrated
individual biases, suggesting a need to combine
all three interests. Combining the three tech-
niques produced similar results to the individual
analyses, showing conservation priorities to be
highest along the east coast. Specifically, the areas
of Port Elizabeth and Durban were ranked high
in all analyses. Consistently, a total of 16 sites
was necessary to represent all species analysed.
Comparisons with similar analyses on fish and
seaweeds revealed similar findings. Existing inver-
tebrate records were shown to be biased towards
centres of high sampling activity, demonstrating
a need of future sampling attention in under-
represented areas.
Key words. Marine invertebrates, South Africa,
endemism, biogeography, marine reserves.
INTRODUCTION
Following the early works of Thompson (1917,
1921, 1924), Clark (1923) and Stephenson (1939,
1944, 1948), much attention has been focused on
documenting the distribution patterns of South
African marine invertebrates (Day, 1967a,b, 1974;
Millard, 1975; Clark & Courtman-Stock, 1976;
Griffiths, 1976; Kensley, 1978, 1981; Gosliner, 1987;
Williams, 1992a,b; Monniot et al. in press). The
wealth of available species records continues to
grow, providing a powerful tool for conservation-
orientated analyses. The overall pattern and trends
of invertebrate distributions have been well described
(Emanuel et al., 1992; Field & Griffiths, 1991),
but direct comparisons between the distributions
of different invertebrate taxa are lacking. The
South African coast is divided into three bio-
geographical provinces, influenced by the flow of
the Benguela upwelling system (west coast), and
the south-flowing Agulhas current (east coast)
(Ekman, 1953; Branch & Branch, 1981). Water
temperatures have the strongest influence on the
biogeographical division into the cool temperate
west coast province, the warm temperate south
coast province, and the subtropical east coast
province (Ekman, 1953; Stephenson & Stephenson,
1972). Conventionally, and for the purposes of
this study, the south coast is delimited by Cape
Point to the west and Port Elizabeth to the east.
A splitting of the east coast into two subprovinces
* Correspondence: GloBallast Programme, International
Maritime Organization, c/o Department of Environmental
Affairs & Tourism, Private Bag X2, Roggebaai 8012, South
Africa. E-mail: Adawad@mcm.wcape.gov.za