NOTE / NOTE
High TNT-transforming activity by a mixed culture
acclimated and maintained on crude-oil-containing
media
Jason T. Popesku, Ajay Singh, Jian-Shen Zhao, Jalal Hawari, and Owen P. Ward
Abstract: A mixed microbial culture originating from a petroleum-contaminated site and maintained on crude oil ex-
hibited high 2,4,6-trinitrotoluene (TNT) transformation activity. Cultivation of the mixed culture in glucose-containing
medium for 29 h resulted in almost complete transformation of 100 ppm TNT. TNT transformation was observed with
both growing and resting cells. With subculturing, it was found that TNT could support growth of the mixed culture
when supplied as sole carbon source, sole nitrogen source, or sole carbon and nitrogen source. The finding that a
mixed microbial culture maintained on crude oil exhibited high TNT transformation activity without prior subculture
on TNT-containing media is novel and may have potential practical applications in the bioremediation of munitions-
contaminated soil and wastewater.
Key words: 2,4,6-trinitrotoluene, mixed culture, transformation, co-metabolism, surfactant.
Résumé : Une culture microbienne mixte provenant d’un site contenant du pétrole et maintenue dans du pétrole brut a
démontré une activité élevée de transformation du 2,4,6-trinitrotoluène (TNT). Une incubation de la culture mixte dans
un milieu renfermant du glucose pendant 29 h a entraîné une transformation quasi complète de 100 ppm de TNT. La
transformation du TNT a été observé avec des cellules en croissance ou au repos. Lors des repiquages, nous avons
constaté que le TNT pouvait soutenir la croissance de la culture mixte lorsqu’il était fourni comme seule source de
carbone, seule source d’azote, ou seule source de carbone et d’azote. Le fait qu’une culture microbienne mixte main-
tenue dans du pétrole brut démontre une activité élevée de transformation du TNT, sans repiquage préalable dans un
milieu contenant du TNT, représente une découverte notoire. Elle aurait des applications pratiques potentielles dans
l’assainissement biologique de sols et d’eaux usées contaminés aux munitions.
Mots clés : 2,4,6-trinitrotoluène, culture mixte, transformation, cométabolisme, surfactant.
[Traduit par la Rédaction] Popesku et al. 366
Barriers to the biodegradation of nitroaromatic com-
pounds include the toxicity of nitroaromatics to microorgan-
isms and the low bioavailability of the compounds because
of their very low solubility. Thus far, a biological pathway
for the complete mineralization of 2,4,6-trinitrotoluene
(TNT) has not been described. Individual aerobic strains of
bacteria converted TNT mainly to various aminonitrotoluenes,
but these compounds could not be further degraded
(Boopathy et al. 1994; Alvarez et al. 1995; Montpas et al.
1997; Hawari et al. 2000; Khachatryan et al. 2000). Some
bacterial strains have been found to grow on TNT as the sole
nitrogen source (French et al. 1998). Several attempts to cul-
ture bacterial strains on TNT as the sole carbon source were
not successful (Boopathy et al. 1994; Khachatryan et al.
2000).
Microbial communities from landfills (Tharakan and
Gordon 1999), anaerobic sludge (Hawari et al. 1998), soil
(Radtke et al. 2000), or lakes (Wikström et al. 2000) con-
taminated with munitions have been shown to be capable of
at least partially degrading TNT. Rates of TNT biotrans-
formation have been shown to increase in soil containing
particulate TNT pretreated with acetone (Radtke et al. 2000)
or in liquid media containing a surfactant, such as Tween 80
(Montpas et al. 1997).
In most of the examples where microbes are used for the
biodegradation of recalcitrant compounds, including the
above-cited cases for TNT, the microorganisms are obtained
from sites containing the contaminant. Consortia or pure cul-
tures are typically isolated by incorporating the contaminant
into selective isolation media, and cultures prior to being
used for contaminant biotransformation or bioremediation
Can. J. Microbiol. 49: 362–366 (2003) doi: 10.1139/W03-049 © 2003 NRC Canada
362
Received 19 February 2003. Revision received 9 June 2003.
Accepted 11 June 2003. Published on the NRC Research
Press Web site at http://cjm.nrc.ca on 17 July 2003.
J.T. Popesku, A. Singh, and O.P. Ward.
1
Department of
Biology, University of Waterloo, Waterloo, ON N2L 3G1,
Canada.
J.-S. Zhao and J. Hawari. Biotechnology Research Institute,
National Research Council, Montréal, QC H4P 2R2, Canada.
1
Corresponding author (e-mail: opward@sciborg.uwaterloo.ca).