1 Metal reduction at low pH by a Desulfosporosinus species: implications for the biological 1 treatment of acidic mine drainage 2 3 John M. Senko 1,2 *, Gengxin Zhang 2§ , Jeffrey T. McDonough 4 , Mary Ann Bruns 3 , William D. 4 Burgos 2 5 1 Department of Geology and Environmental Science, The University of Akron; 2 Department of 6 Civil and Environmental Engineering and 3 Department of Crop and Soil Sciences, The 7 Pennsylvania State University; 4 ARCADIS U.S. Inc. 8 9 10 *Author to whom correspondence should be addressed: 11 John M. Senko 12 Department of Geology and Environmental Science 13 216 Crouse Hall 14 The University of Akron 15 Akron, OH 44325 16 Phone: 330-972-8047 17 Fax: 330-972-7611 18 Email: senko@uakron.edu 19 § Current address: Environmental Science Division, Oak Ridge National Laboratory 20 Running head: Metal and sulfate reduction at low pH 21 22