For Peer Review Only               ! "   #$%& $ " " ’ % ( ) " *+ ,%%-  ) ! & ." " /(0+ % # 1  2%+ - 3* 3)+ - 2" 14+ % ) $+ & & "& "" 5$*+ .&&- 2" 14+ "!+ 6! & 4  + "&- 2" 14+ "!+ 6! & 4   %%+ .- 2" 14+ "!+ 6! & 4  73& &$+ $+ %"% 4+  "" E-mail: cavanagh@metronet.co.uk URL: http://mc.manuscriptcentral.com/cavp Avian Pathology peer-00687801, version 1 - 15 Apr 2012 Author manuscript, published in "Avian Pathology 40, 02 (2011) 139-144" DOI : 10.1080/03079457.2010.543414