ORIGINAL ARTICLE
Aromatase expression in human peripheral blood
leucocytes (PBLs) and in various tissues in primates: studies
in elderly humans and cynomolgus monkeys
Elisa Pignatti
1,2
, Livio Casarini
1,2
, Sara Scaltriti
1
, Joachim Wistuba
3
, Stefan Schlatt
3
, Alessandra
Rossi
1
, Asmaa Lachhab
1
, Erica Taliani
1
, Cesare Carani
1
& Manuela Simoni
1,2
1 Department of Biomedical, Metabolic and Neural Sciences, University of Modena and Reggio Emilia Modena, Italy
2 Center of Genome Research (CGR), University of Modena and Reggio Emilia Modena, Italy
3 Institute of Reproductive and Regenerative Biology, Centre of Reproductive Medicine and Andrology, University Mu ¨ nster Mu ¨ nster, Germany
Keywords
ageing – gene expression – oestrogens –
protein production
Correspondence
Manuela Simoni, Department of Biomedical,
Metabolic and Neural Sciences, University
of Modena and Reggio Emilia, NOCSAE, via
Giardini 1355, 41126 Baggiovara, Modena,
Italy.
Tel.: +39 059 3961800;
fax: +39 059 3961335;
e-mail: manuela.simoni@unimore.it
Accepted July 30, 2012.
Abstract
Background Previous analysis of aromatase gene and protein expression in
peripheral blood leucocytes (PBLs), studied in children and adults, was
extended to elderly subjects. In addition, we assessed whether aromatase
expression in PBLs could be used as a parameter of aromatase expression in
other tissues, using the cynomolgus monkey as model.
Methods Real-time PCR analysis of aromatase gene expression and protein
evaluation by Western blot was performed in PBLs of human elderly subjects
and in various tissues from cynomolgus monkeys.
Results No gender-related difference in CYP19A1 mRNA and protein
expression in PBLs from human elderly women and men was found. In
elderly male cynomolgus monkeys, CYP19A1 mRNA and protein were
expressed in all cells and tissues analysed, with the lowest levels in PBLs but
no clear-cut correlation with other tissues.
Conclusions Aromatase expression in PBLs in elderly human subjects is not
gender-related and cannot be a surrogate of aromatase expression for other
tissues.
Introduction
Aromatase, a member of cytochrome P450 superfamily,
is the key enzyme in vertebrates involved in the conver-
sion of testosterone to oestradiol and androstenedione
to estrone [28]. The enzyme consists of a haem group
and a polypeptide chain of 513 amino-acid residues
derived from the CYP19A1 gene localized on chromo-
some 15q21.1 [9]. Aromatase is expressed in a large
number of cells and tissues [8] in a tissue-specific man-
ner, regulated by different promoter regions and by an
incompletely understood alternative splicing mechanism
[11, 27]. Several studies investigating aromatase expres-
sion and/or activity suggest that oestrogen imbalance
may result in pathological conditions such as breast
cancer, infertility, reproductive defects, metabolic
disorders and abnormalities in bone development [8].
Therefore, knowing the physiological and pathological
expression and activity of aromatase by tissue, age and
gender is of utmost clinical significance. Such studies,
however, are hampered by the limitations in obtaining
normal human tissue from different organs.
Following the identification of a novel exon 1
sequence and its aberrant expression in mammary
gland, in fibroblasts and in lymphocytes of patients with
aromatase excess syndrome [31], a possible correlation
between aromatase in target tissues and in peripheral
blood leucocytes (PBLs) was suggested, prompting the
study of aromatase in PBLs obtained from young adult
women and men and prepubertal children [35]. The
comparison between PBL aromatase mRNA and pro-
tein level in different ages during the follicular and luteal
J Med Primatol 41 (2012) 372–383
© 2012 John Wiley & Sons A/S 372
J Med Primatol doi:10.1111/jmp.12000