J. Basic Microbiol. 44 (2004) 1, 10 – 16 DOI: 10.1002/jobm.200310281
© 2004 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim 0233-111X/04/0102-0010
(Department of Food Science, University of Foggia, via Napoli, 25, 71100 Foggia, Italy
Molecular characterization of lactic acid populations
associated with wine spoilage
L. BENEDUCE, G. SPANO*, A. VERNILE, D. TARANTINO and S. MASSA
(Received 04 April 2003/Accepted 01 July 2003)
We have investigated the prevalence of spoilage lactic acid bacteria (LAB) in table wines produced in
the Apulia region. The occurrence of LAB was evaluated in wines produced with low sulphur dioxide
doses and not supplemented with selected malolactic starters such as Oenococcus oeni. About 150
strains were isolated from wine must and a molecular characterization was performed using PCR-
based techniques. Most of the strains analysed belonged to Lactobacillus plantarum species. However,
some of the strains were identified as Pediococcus damnosus and Leuconostoc sp. The amplified
fragments of Pediococcus damnosus were cloned and sequenced. The coding sequence was highly
homologous to that of the ropy plasmid confirming that the isolated strain was a ropy
+
Pediococcus
damnosus. In all the samples analysed, the final must pH value reached was relatively high (from 3.78
to 3.90). The high pH values had probably negatively influenced (counteracted) the activity of sulphur
dioxide added, allowing proliferation of spoilage wine microorganisms.
During malolactic fermentation (MLF), lactic acid bacteria (LAB) lead to a biological
deacidification of wine, due to the conversion of bicarboxylic malic acid into monocarb-
oxylic lactic acid and a broad range of secondary compositional modifications of great im-
portance for taste and flavour improvement of wine (LIU 2002). When all the malic acid is
degraded, wines are usually stabilized by sulfiting. Sulphur dioxide is at this stage of wine-
making the only authorized and efficient agent for the stabilization of wine (LONVAUD-
FUNEL 1999). Most of the bacteria and possibly the remaining yeasts are sensitive to sulfur
dioxide. However, for health reasons, high doses of sulphur dioxide are not recommended
(DELFINI et al. 1992). Furthermore, the activity of sulphur dioxide is related to pH. In wines
with high pH, the ineffectiveness of sulphur dioxide may enhance survival of spoilage bac-
teria. In these conditions, Lactobacillus and Pediococcus are frequently predominant and
can induce spoilage. Only physical treatments such as heat treatment or filtration can even-
tually eliminate all the viable bacteria (LONVAUD-FUNEL 1999).
In some areas, especially in south of Italy, although the majority of wine makers still
prefer to produce homemade wines, some of them has begun to build up small commercial
wineries. In either case, however, no selected malolactic starters are used and the table
wines produced sometimes have, a low quality and bitter taste. Furthermore, low doses of
sulphur dioxide are usually added during or at the end of MLF.
Understanding the importance of using selected starters during MLF and identifying the
predominant LAB population during MLF should be of great value in order to predict either
the frequency of spoilage microorganisms or to improve wine quality produced by small
wineries. The improvement of molecular tools, usually based on PCR techniques, has al-
lowed a fast and sensitive characterization of the majority of wine LAB. However, the same
tools should be available and easily understandable even for small winery that might use
them to improve their wine quality.
* Corresponding author: Dr. G. SPANO; e-mail: g.spano@unifg.it