pubs.acs.org/JAFC Published on Web 09/10/2009 © 2009 American Chemical Society
J. Agric. Food Chem. 2009, 57, 9427–9436 9427
DOI:10.1021/jf901880p
Antioxidant Capacity and Lipid Characterization of Six
Georgia-Grown Pomegranate Cultivars
GARIMA PANDE AND CASIMIR C. AKOH*
Department of Food Science and Technology, University of Georgia, Athens, Georgia 30602-2610
Six pomegranate (Punica granatum) cultivars were investigated for their antioxidant capacity and lipid
profile. Total polyphenols were determined according to the Folin-Ciocalteau method. Major organic
acids and phenolic compounds were analyzed by RP-HPLC. Two in vitro antioxidant assays, ferric
reducing antioxidant power and Trolox equivalent antioxidant capacity, were used to assess anti-
oxidant capacity. Total lipid was extracted according to the Folch method, and fatty acid methyl esters
were determined by GC. Tocopherols and phospholipids were identified and quantified by NP-HPLC
using a fluorescence detector for tocopherols and an evaporative light scattering detector for
phospholipid analysis. Phytosterols were analyzed by GC. The predominant organic acid was citric
acid followed by malic acid. The peel fraction had the highest total hydrolyzable tannins content
(4792.3-6894.8 mg/100 g of FW). Overall, the highest antioxidant capacity was found in leaves
followed by peel, pulp, and seed. Pomegranate seed had an average lipid content of 19.2% with
punicic acid as the predominant fatty acid. Pomegranate seed had high contents of R-tocopherol
(161.2-170.1 mg/100 g) and γ-tocopherol (80.2-92.8 mg/100 g).
KEYWORDS: Antioxidant capacity; fatty acids; organic acids; phospholipids; phytosterols; polyphenols;
Punica granatum; tocopherols
INTRODUCTION
Phytochemicals, particularly antioxidants from natural sources
such as fruits and vegetables, have gained popularity because
many epidemiological studies have shown their protective proper-
ties against several chronic diseases such as cancer and cardio-
vascular diseases ( 1 ). Products and intermediates of oxidative
stress pathways are associated with several chronic diseases.
Oxidative stress is the disturbance of the pro-oxidant-antioxidant
balance in favor of the former, leading to potential damage ( 2 ).
Interest in natural antioxidants has increased during the past few
decades because of the adverse effects shown by synthetic anti-
oxidants and also due to the worldwide trend to avoid or minimize
the use of artificial food additives ( 3 ). The most abundant anti-
oxidants in fruits are polyphenols, vitamins, and carotenoids.
The lipid profiles of several fruits and their seeds have been
characterized and various bioactive compounds isolated ( 4 , 5 ).
Determination of the lipid classes of the sample may aid in decid-
ing its application in food, health, and other industries. This in
turn will help in identifying the potential of the crop as a main-
stream agricultural product.
Pomegranate (Punica granatum), belonging to family Punica-
ceae, has been used in several traditional medicine systems. It is a
rich source of various bioactive compounds demonstrating anti-
oxidant and anti-inflammatory activities ( 6 ). Pomegranate seed
oil comprising 12-20% of the total seed weight consists of
approximately 80% conjugated octadecatrienoic fatty acids,
mainly punicic acid ( 6 ). Highest antioxidant activity has been
shown in pomegranate peel compared to pulp and seed frac-
tions ( 7 ), which can be attributed to its high content of tannins,
especially punicalagin isomers. Pomegranate juice is bright red in
color due to its high content of flavonoids and anthocyanins.
Pomegranate juice exhibits antiatherogenic activities ( 8 ). Leaves,
flowers, bark, and roots also contain distinctive compounds
having potent physiological effects.
Pomegranate is a minor fruit crop of Georgia, but with the
current commercial and nutritional potentials, a detailed study
may help in its cultivar selection and application. The peels and
seeds, which are usually disposed of as waste material in many
food-processing industries, could be a rich source of beneficial
phytochemicals. From economic and environment points of view
disposing of such wastes should be avoided. Recovery of bioac-
tive compounds would help to make the recycling of wastes
economically viable and also result in value addition to these
minor crops. The main objective of this research was to compare
six Georgia-grown pomegranate cultivars in terms of their
organic acid contents, antioxidant capacities, and lipid profiles.
MATERIALS AND METHODS
Plant Material. Six pomegranate (P. granatum, Punicaceae) cultivars,
R19, R26, Cvg-Eve, North, Crab, and Cranberry, along with their leaves
were obtained from Ponder farm, a University of Georgia operated farm
near Tifton, GA. Managed plant growth conditions were minimal.
Irrigation was not performed, and only natural rainfall was utilized. No
supplemental fertilizers were applied. They were planted in a loamy-sand
soil (sand, 86%; silt, 7%; and clay, 7%). Some pruning was performed, but
they were not managed in a commercial manner. They were planted from
1990 to 1993 and have been neglected until spring of 2008, when some
*Author to whom correspondence should be addressed [telephone
(706) 542-1067; fax (706) 542-1050; e-mail cakoh@uga.edu].
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Published on September 10, 2009 on http://pubs.acs.org | doi: 10.1021/jf901880p