Soil Environ. 31(1): 30-36, 2012
www.se.org.pk
Online ISSN: 2075-1141
Print ISSN: 2074-9546
*Email: kannikamsu13@yahoo.com
© 2012, Soil Science Society of Pakistan (http://www.sss-pakistan.org)
Selection of efficient salt-tolerant bacteria containing ACC deaminase for
promotion of tomato growth under salinity stress
Kannika Chookietwattana
*
and Kedsukon Maneewan
Department of Biotechnology, Faculty of Technology, Mahasarakham University, Thailand
Abstract
For successful application of plant growth promoting bacteria (PGPB) in salt-affected soil, bioinoculant with salt-
tolerant property is required in order to provide better survival and perform well in the field. The present study
aimed to select the most efficient salt-tolerant bacterium containing 1-aminocyclopropane-1-carboxylic acid (ACC)
deaminase from eighty four bacterial strains and to investigate the effects of the selected bacterium on the
germination and growth of tomato (Licopersicon esculentum Mill. cv. Seeda) under saline conditions. The Bacillus
licheniformis B2r was selected for its ability to utilize ACC as a sole nitrogen source under salinity stress. It also
showed a high ACC deaminase activity at 0.6 M NaCl salinity. Tomato plants inoculated with the selected
bacterium under various saline conditions (0, 30, 60, 90 and 120 mM NaCl) revealed a significant increase in the
germination percentage, germination index, root length, and seedling dry weight especially at salinity levels ranging
from 30-90 mM NaCl. The work described in this report is an important step in developing an efficient salt-tolerant
bioinoculant to facilitate plant growth in saline soil.
Key words: ACC deaminase, salt stress-induced ethylene, salt-tolerant bacteria
Introduction
Salt-affected soil can result in either the reduction of
crop productivity or abandonment of many agricultural
areas (Maas, 1990). Under high soil salinity, plant
biosynthesizes ethylene by using 1-aminocyclopropane-1-
carboxylic acid (ACC) as a precursor, and hence called salt
stress-induced ethylene. Numerous reports have
documented that ethylene could inhibit the elongation of
plant root and shoot (Jusaitis, 1986; Penrose and Glick,
2003), suppress leaf expansion (Peterson et al., 1991) and
promote epinasty (Abeles et al., 1992). Thus, reduction of
salt stress-induced ethylene production in plant is one of the
most crucial strategies to increase agricultural production in
saline soil.
In the last decade, the concept of plant growth
promoting bacteria (PGPB) containing ACC-deaminase for
promotion of plant growth under environmental stress
conditions has gained importance (Berg, 2009). These
bacteria are capable of metabolizing ACC in the root of
developing plants, thereby reducing the adverse effects of
ethylene on plant growth (Ghosh et al., 2003; Glick, 2005;
Hontzeas et al., 2006). However, the ability of inoculated
bacteria to survive, outcompete with the native microflora,
and colonize in the rhizosphere remains to be the challenge
for successful application (Bashan, 1998) especially in
saline soil since high salinity could influence the survival,
growth and activity of microorganisms (Bremer and
Krämer, 2000). The salt-tolerant ACC deaminase-
containing bacteria could thus be advantageous over others
to thrive in a new saline environment in the sufficient
numbers to deliver beneficial effects on plants. Although
the achievements of ACC deaminase-containing bacteria in
promoting plant growth under various environmental
stresses were reported immensely (Grichko and Glick,
2001; Dey et al., 2004; Mayak et al., 2004; Kausar and
Shahzad, 2006; Shaharoona et al., 2006; Belimov et al.,
2008; Zahir et al., 2008), little information is available on
the effectiveness of salt-tolerant bacteria containing ACC
deaminase under high salinity (Nadeem et al., 2006).
Therefore, in this research study, attempts have been made
to screen and select an efficient salt-tolerant ACC
deaminase containing bacterium strain which could utilize
ACC as a sole nitrogen source under salt- stressed
conditions. Its effectiveness for growth promotion of
tomato (Licopersicon esculentum Mill cv. Seeda) under in
vitro and saline conditions has also been evaluated.
Materials and Methods
Microorganisms and culture conditions
Eighty-four bacterial strains, which were isolated from
saline soil in Thailand, and identified on the basis of
phenotypic characterization, and 16S rDNA sequence
analysis from the previous work of Chookietwattana
(2003), were used in the experiment. For inoculum
preparation, bacterial strains were individually cultured at