P. ostreatus Peroxidase Profiles 415
Applied Biochemistry and Biotechnology Vol. 102–103, 2002 415
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Mn
2+
Alters Peroxidase Profiles
and Lignin Degradation
by the White-Rot Fungus Pleurotus ostreatus
Under Different Nutritional
and Growth Conditions
RONI COHEN, LIMOR PERSKY, ZAHIT HAZAN-EITAN,
ODED YARDEN, AND YITZHAK HADAR*
Department of Plant Pathology and Microbiology
and The Otto Warburg Center for Biotechnology in Agriculture,
Faculty of Agricultural, Food and Environmental Quality Sciences,
The Hebrew University of Jerusalem, PO Box 12, Rehovot 76100, Israel,
E-mail: hadar@agri.huji.ac.il
Abstract
The white-rot fungus Pleurotus ostreatus produces two types of extracellu-
lar peroxidases: manganese-dependent peroxidase (MnP) and versatile per-
oxidase (VP). The effect of Mn
2+
on fungal growth, peroxidase activity
profiles, and lignin degradation by P. ostreatus was studied in liquid culture
and under solid-state fermentation conditions on perlite, the latter resem-
bling the natural growth conditions of this fungus. The fungus was grown in
either a defined asparagine-containing basidiomycete selective medium
(BSM) or in a rich peptone medium (PM). Biomass production, as deter-
mined by respiration experiments in solid-state fermentation and liquid
cultures and fungal growth on Petri dishes, was higher in the PM than in
the BSM. Mn
2+
affected biomass production only in the PM on Petri dishes.
In the nonamended PM, high levels of MnP and VP activity were detected
relative to the nonamended BSM. Nevertheless, a higher rate of
14
C-lignin
mineralization was measured in the Mn
2+
-amended BSM, as determined
during the course of 47 d of fermentation. Mn
2+
amendment of the PM
increased mineralization rate to that obtained in the Mn
2+
-amended BSM.
The enzyme activity profiles of MnP and VP were studied in the BSM using
anion-exchange chromatography. In the nonamended BSM, only minute
levels of MnP and VP were detected. On Mn
2+
amendment, two MnP isoen-
zymes (B1 and B2) appeared. Isoenzyme B2 was purified and showed 100%
identity with the MnP isoenzyme purified in our previous study from
PM-solid-state fermentation (P6). P6 was found to be the dominant isoenzyme