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Heritability analyses show visit-to-visit blood pressure
variability reflects different pathological phenotypes in
younger and older adults: evidence from UK twins
Cristina Menni
a
, Massimo Mangino
a
, Feng Zhang
a
, Gail Clement
a
, Harold Snieder
b
,
Sandosh Padmanabhan
c
, and Tim D. Spector
a
Background: Clinic and long-term average blood pressure
(BP) are heritable traits with estimates of heritability
ranging from 0.31 to 0.68. Long-term visit-to-visit BP
variability (BPV) is emerging as a new cardiovascular risk
predictor, though it is unclear if this is completely
independent of BP. We hypothesize that BPV should
demonstrate the same pattern of additive genetic, shared
environmental and unique environmental variance as BP, if
both are phenotypic surrogates.
Method: We studied 2889 twin pairs not on any BP-
lowering therapy from the Twins UK cohort, and estimated
the additive genetic variance for baseline BP, long-term
average BP, BP trajectory (rate of change of BP in mmHg/
year) and BPV (coefficient of variation and average real
variability over an average of 3.2 visits). Heritability
estimates were obtained by structural equation modelling
adjusting for age, age
2
, sex and BMI.
Results: The heritabilities for baseline SBP and DBP were
0.51 (95% confidence interval 0.49, 0.53) and 0.56 (0.54,
0.58); long-term average SBP and DBP were 0.56 (0.53,
0.59) and 0.61 (0.58, 0.64); and systolic and diastolic
trajectories over 10 years were 0.49 (0.46, 0.52) and 0.29
(0.27, 0.32), respectively. Both overall systolic and diastolic
BPV showed no additive genetic variance contributing to
the phenotypic variation, but after stratification by age, the
younger subgroup (<51 years) showed heritability
estimates of 0.44 (0.38, 0.50) for coefficient of variation
and 0.35 (0.29, 0.41) for average real variability.
Conclusion: Age is a major factor that influences
heritability estimation of BPV and it is likely that BPV in
younger and older age groups may reflect different
pathological phenotypes.
Keywords: blood pressure variability, heritability, twins
Abbreviations: A, additive genetic variance; AIC, Akaike
information criterion; AVR, average real variability; BP,
blood pressure; BPV, blood pressure variability; C, shared/
common environmental variance; CKD, chronic kidney
disease; E, unique environmental variance; h
2
, heritability
INTRODUCTION
L
ong-term visit-to-visit blood pressure variability
(BPV) is emerging as an independent cardiovascular
risk predictor from post-hoc analysis of large clinical
trials and population cohorts [1 – 3]. It is well established that
blood pressure (BP) is heritable with estimates of herit-
ability ranging from 0.31 to 0.68 [4–8], and high BP is an
independent predictor of early mortality [9]. It is unclear
whether the predictive value of long-term BPV is independ-
ent of clinic or ambulatory BP readings, though there is
some evidence that the risk posed by BPV is independent of
average BP [1,10]. As BPV correlates with clinic and long-
term average BP [3,11], we hypothesize that it should also
demonstrate the same pattern of additive genetic (A),
shared environmental (C) and unique environmental (E)
variance, if it is a phenotypic surrogate of BP. Recently,
heritability analysis of short-term BPV from ambulatory BP
data showed daytime BPV was not heritable, whereas
night-time BPV was 0.33–0.36 heritable [12]. We analysed
the heritabilities of BP, long-term average BP, BP trajectory
and BPV in a cohort of adult twins to determine if BPV is
independent of BP.
METHODS
Study population
Study participants were twins enrolled in the Twins UK
registry, a national register of adult twins recruited as
volunteers by successive media campaigns without select-
ing for particular diseases or traits [13]. The registry
Journal of Hypertension 2013, 31:000–000
a
Department of Twin Research & Genetic Epidemiology, King’s College London,
London, UK,
b
Department of Epidemiology, University of Groningen, University
Medical Center Groningen, The Netherlands and
c
British Heart Foundation (BHF)
Glasgow Cardiovascular Research Centre, University of Glasgow, Glasgow, UK
Correspondence to Dr Sandosh Padmanabhan, BHF Glasgow Cardiovascular Research
Centre, Institute of Cardiovascular and Medical Sciences, University of Glasgow,
Glasgow G12 8TA, UK. E-mail: sandosh.padmanabhan@glasgow.ac.uk or Prof Tim D
Spector, Department of Twin Research and Genetic Epidemiology, King’s College
London, St. Thomas’ Hospital, London SE1 7EH, UK. Tel: +4420 7188 5555; fax: +44
20 7188 6761; e-mail: tim.spector@kcl.ac.uk
Received 4 December 2012 Revised 21 May 2013 Accepted 16 July 2013
J Hypertens 31:000–000 ß 2013 Wolters Kluwer Health | Lippincott Williams &
Wilkins.
DOI:10.1097/HJH.0b013e32836523c1
Journal of Hypertension www.jhypertension.com 1
Original Article