Hindawi Publishing Corporation
Evidence-Based Complementary and Alternative Medicine
Volume 2013, Article ID 261978, 6 pages
http://dx.doi.org/10.1155/2013/261978
Research Article
Dodonaea viscosa var. angustifolia Inhibits Germ Tube and
Biofilm Formation by C. albicans
Serisha Devi Naicker and Mrudula Patel
Division of Oral Microbiology, Department of Clinical Microbiology and Infectious Diseases, National Health Laboratory Services and
Faculty of Health Sciences, University of the Witwatersrand, Private Bag 3, Wits, Johannesburg 2050, Gauteng, South Africa
Correspondence should be addressed to Mrudula Patel; mrudula.patel@wits.ac.za
Received 26 July 2013; Accepted 5 September 2013
Academic Editor: Veronique Seidel
Copyright © 2013 S. D. Naicker and M. Patel. his is an open access article distributed under the Creative Commons Attribution
License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly
cited.
he virulence factors of Candida albicans are germ tube and bioilm formation, adherence to host tissues, and production of
hydrolytic enzymes. his study investigated the efect of Dodonaea viscosa var. angustifolia extract on the germ tube and bioilm
formation of C. albicans. Serum containing the three subinhibitory concentrations of leaf extract was inoculated with C. albicans,
incubated, and viewed under a light microscope. Number of cells with germ tube was recorded and the results were analysed
using Schefe test for pairwise comparison. Bioilms were grown on coverslips in the presence of plant extracts and processed
for scanning electron microscopy (SEM). Planktonic cells were grown in the presence of plant extract for 6h and processed for
electron microscopy (TEM). he crude plant extract signiicantly ( < 0.01) reduced the germ tube formation of C. albicans at 3.125
(85.36%), 1.56 (61.91%), and 0.78mg/mL (26.27%) showing a concentration dependent efect. SEM results showed concentration
dependent reduction in bioilm and hyphae formation. TEM results showed that the plant extract caused damage to the cell wall and
cell membrane. DVA extract has ability to reduce virulence of C. albicans by inhibiting germ tube and bioilm formation through
damage to the cell wall. herefore, it has therapeutic potential.
1. Introduction
Oral candidiasis is an opportunistic infection of the oral cav-
ity which usually occurs in the elderly and immunocompro-
mised individuals [1]. Candida albicans is the most common
species of yeast isolated from patients with oral candidiasis
[2]. It can cause supericial infections such as thrush and
denture stomatitis; however in more severe cases, it causes
life-threatening systemic mycoses. Most of these commensal
C. albicans cells proliferate as budding yeasts. However, when
the epithelial barrier is breached due to trauma, immuno-
suppression, or hormonal alterations, the budding form is
converted into a hyphal form which causes invasion of sub-
mucosal tissues [3]. he ability to switch between yeast and
hyphal morphologies can play a role in the virulence of C.
albicans [4]. Other virulence properties are adherence to host
tissues and prostheses and ability to produce hydrolytic enzy-
mes. Some of these factors are interrelated. C. albicans cells
bearing germ tubes are more adherent to buccal epithelial
cells (BEC) than yeast forms of C. albicans [5] due to the anti-
gens found on germ tubes [6].
Many efective antifungal agents such as amphotericin B,
nystatin, and luconazole are available. However, poor solu-
bility, stability, and absorption, toxicity, and the development
of drug resistance are a problem with existing drugs [7].
A novel approach to therapy has been suggested by some
researchers, where the virulence of C. albicans can be targeted
instead of an antimicrobial property. his would be ideal
where the causative organism is a part of the normal lora,
such as C. albicans in the oral cavity [2, 8]. Pathogenic chara-
cteristics such as germ tube and bioilm formation and pro-
duction of tissue damaging enzymes are possible targets of
new drugs.
Many medicinal plants used in Africa have been explored
for their anticandida activities [9]. D. viscosa var. angustifolia
(DVA) belongs to the Sapindaceae family and it is found in
many parts of the world including South Africa. Leaves and
tips of the twigs have many medicinal properties and they
are traditionally used to treat colds, fever, lu, sore throats,
and oral thrush worldwide [10, 11]. It has demonstrated in
vitro anticandida activity at high (MIC of 6.25–25mg/mL)