MYERS, C. W., AND W. E. DUELLMAN. 1982. A new species of Hyla from Cerro Colorado, and other treefrog records and geographical notes from western Panama. American Museum Novitates 2752:1–32. RICHARDS, S. J., AND D. T. ISKANDAR. 2001. A new species of treefrog (Anura, Hylidae, Litoria) from the mountains of Irian Jaya, Indonesia. Alytes 18:141–152. SAVAGE, J. M., AND W. R. HEYER. 1967. Variation and distribution in the treefrog genus Phyllomedusa in Costa Rica, Central America. Beitra ¨ge zur Neotropischen Fauna 5:111–131. TYLER, M. J. 1962. New hylid frog from the central highlands of New Guinea. Records of the South Australian Museum 14:253–257. ———. 1968. Papuan hylid frogs of the genus Hyla. Zoologische Verhandelingen 96:1–203. TYLER, M. J., AND M. DAVIES. 1978. Species-groups within the Australopapuan hylid frog genus Litoria Tschudi. Australian Journal of Zoology, Supplemental Series 63:1–47. Accepted: 16 July 2003 Associate Editor: Chris Sheil APPENDIX I Additional Specimens Examined Litoria nigropunctata.—Papua New Guinea: Madang Province: Binek, near Madang, UPNG 2330–34; Morobe Province: Lae, UPNG 3649–52, 3658. Herpetologica, 60(1), 2004, 103–116 Ó 2004 by The Herpetologists’ League, Inc. A NEW SPECIES OF ELEUTHERODACTYLUS (ANURA: LEPTODACTYLIDAE) FROM THE NORTHWESTERN LOWLANDS OF ECUADOR JUAN M. GUAYASAMIN 1 Natural History Museum and Biodiversity Research Center, Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045-7561, USA ABSTRACT: A new species of Eleutherodactylus is described from the lowlands of northwestern Ecuador. The new species is a small frog (snout–vent length ,23 mm) characterized by: expanded and spadate disc cover on some fingers (usually III and IV) and most toes (usually III–V); bifid palmar tubercle; and brown dorsum with dark brown W-shaped scapular mark. A detailed osteological description of the new species is provided. Some of the osteological characters include relatively large nasals, small sphenethmoid, eight procoelous presacral vertebrae, manus with six carpal elements, and pes with four tarsal elements. A small portion of the frontoparietal fontanelle is exposed between the frontoparietals. Although the new species is small, there is no loss of skull bones and phalangeal elements. Key words: Ecuador; Eleutherodactylus; Leptodactylidae; New species; Osteology THE ANURAN genus Eleutherodactylus oc- curs in Central and South America, as well as the West Indies, and comprises more than 600 recognized species (Duellman, 1993; Frost, 2002). In Ecuador, there are 136 described species of Eleutherodactylus, which is the most diverse and usually the most abundant anuran genus in the cloud forests and adjacent lowland rainforests (Coloma and Quiguango, 2000– 2002; Lynch and Duellman, 1997). Although knowledge of the alpha-level taxonomy of Eleutherodactylus in Ecuador has increased considerably in recent years (e.g., Lynch and Duellman, 1995, 1997; Duellman and Pramuk, 1999), undescribed species continue to be discovered. The latter is especially true in western Ecuador, where the discovery of at least five new species was announced (Lynch and Duellman, 1997:190–191). Herein, I de- scribe a new species of Eleutherodactylus from the lowlands of northwestern Ecuador and include a detailed description of the cranial and postcranial osteology of the species. 1 CORRESPONDENCE: e-mail, juanm@ku.edu March 2004] HERPETOLOGICA 103