SHORT COMMUNICATION
Modulation of P-glycoprotein ATPase Activity
by Some Phytoconstituents
I. A. Najar,
1
B. S. Sachin,
1
S. C. Sharma,
1
N. K. Satti,
2
K. A. Suri
2
and R. K. Johri
1
*
1
Division of Pharmacology, Indian Institute of Integrative Medicine, (Formerly Regional Research Laboratory), CSIR, Jammu 180
001, India
2
Division of Natural Products Chemistry, Indian Institute of Integrative Medicine, (Formerly Regional Research Laboratory),
CSIR, Jammu 180 001, India
In the present investigation 16 phytoconstituents, which are active moieties found in several medicinal herbs,
have been evaluated for their P-glycoprotein (P-gp) stimulation/inhibition profiles using a P-gp-dependent
ATPase assay in rat jejunal membrane (in vitro). Acteoside, agnuside, catechin, chlorogenic acid, picroside -II
and santonin showed an inhibitory effect. Negundoside, picroside -I and oleanolic acid caused a stimulatory
effect. Andrographolide, apocyanin, berberine, glycyrrhizin, magniferin and piperine produced a biphasic
response (stimulation at low concentration and inhibition at high concentration). The results suggested that a
possible interaction of these phytoconstituents at the level of P-gp, could be an important parameter in deter-
mining their role in several key pharmacodynamic events. Copyright © 2009 John Wiley & Sons, Ltd.
Keywords: P-glycoprotein; ATPase activity; phytoconstituents.
INTRODUCTION
P-glycoprotein (P-gp) is a membrane-located protein
of the ABC superfamily found primarily in drug-
eliminating organs. It is recognized as an important host
defense mechanism mitigating the cellular burden of
exogenous and endogenous substrates. P-gp is a poly-
specific efflux transporter that can pump out a very
broad range of substrates. In addition, the role of this
protein in the regulation of oral bioavailability of several
clinically important drugs, and in potential drug–drug
interactions has also drawn considerable attention
(Marchetti et al., 2007).
In recent times an enormous increase in the consump-
tion of herbal remedies along with prescription drugs
has been witnessed. Most of such remedies are derived
from a variety of medicinal plants. The majority of such
herbal medicinal products contain a wide array of bioac-
tive compounds which have been shown to modulate
P-gp function (Deferme and Augustijns, 2003; Brand
et al., 2006). This has brought into focus a substantial
impact of P-gp on herb–drug interactions of wide clini-
cal relevance (Izzo, 2004).
P-gp is an unusual ATP-driven transporter, in that it
has a low affinity for ATP and exhibits a high level of
constitutive or basal ATPase activity. This protein has
12 transmembrane domains contained in two homolo-
gous halves and two ATP-binding cassette domains in
each half that catalyse ATP hydrolysis (Kokubu et al.,
1997). One characteristic of this protein is that it couples
binding and hydrolysis of ATP at the two nucleotide-
binding domains to drug export by transmembrane
domains (Tombline et al., 2008). Therefore ATP hydro-
lysis happens to be an inherent property of P-gp.
However, no evaluation of the effect of most herbal
moieties on P-gp ATPase activity had been undertaken
prior to a current investigation of which the results on
some phytoconstituents presented herein are a part.
MATERIALS AND METHODS
Chemicals. ATP, dithiotreitol (DTT), ethylene-bis
(oxyethylenenitrilo) tetracetic acid (EGTA), Hepes,
ouabain octahydrate, sodium ortho-vanadate, Tris-HCl
were purchased from Sigma Chemical Co. (St Louis,
MO, USA). All other chemicals and reagents were of
high purity analytical grade.
Animals. Healthy Wistar rats (150–175 g body weight)
of both sexes were maintained in regulated environ-
mental conditions (well-ventilated with >10 air
changes/h; 12 h light/dark photoperiod; temperature
28 ± 2 ºC; relative humidity, 60 ± 10%), according
to CPCSEA guidelines. Animal experiments were
approved by the Institutional Animal Ethics Committee
and were performed as per the Guidelines for Animal
Care as recommended by the Indian National Academy,
New Delhi (1992). Animals were fed with standard
pelleted diet (Ashirwad Industries, Chandigarh, India)
and water was provided ad libitum. Seven days after
acclimatization, the animals were used.
* Correspondence to: Dr R. K. Johri, Division of Pharmacology, Indian
Institute of Integrative Medicine, (Formerly Regional Research Labora-
tory), CSIR, Jammu 180 001, India.
E-mail: rakeshkjohri@rediffmail.com; rkjohri@iiim.res.in
Received 20 May 2009
Revised 27 May 2009
Copyright © 2009 John Wiley & Sons, Ltd. Accepted 28 May 2009
PHYTOTHERAPY RESEARCH
Phytother. Res. 24: 454–458 (2010)
Published online 3 August 2009 in Wiley InterScience
(www.interscience.wiley.com) DOI: 10.1002/ptr.2951