Journal of Microbiology Research 2014, 4(4): 161-169
DOI: 10.5923/j.microbiology .20140404.02
Toxicity of Binary Mixtures of Formulated Glyphosate
and Phenols to Rhizobium Species Dehydrogenase Activity
C. O. Nweke
1,*
, N. C. Ahumibe
2
, J. C. Orji
1
1
Department of Microbiology, Federal University of Technology, P.M.B. 1526, Owerri, Nigeria
2
Department of Biotechnology, Federal University of Technology, P.M.B. 1526, Owerri, Nigeria
Abstract Acute toxicities of formulated glyphosate (Roundup®) in binary mixtures with 2,4-dichlorophenol,
4-chlorophenol and phenol were determined based on inhibition of 2-(p-Iodophenyl)-3-(p-nitrophenyl)-5-phenyl tetrazolium
chloride (INT)-dehydrogenase activity in Rhizobium species. The phenolic compound: glyphosate mixture ratios (%) were
100:0, 20:80, 50:50, 60:40, 20:80 and 0:100 for the respective mixtures in the concentration range of 0 – 5000 mg/l. The
effective doses (IC
p
s) were estimated using hormetic dose-response model. The median inhibitory concentrations (IC
50
) of
the formulated glyphosate, phenol, 4-chlorophenol and 2,4-dichlorophenol were 661.614 ± 33.234 mg/l, 1064.190 ± 87.286
mg/l, 158.884 ± 8.185 mg/l and 46.378 ± 2.504 mg/l respectively. The joint action toxicity of the mixtures on test organism
was evaluated with isobolographic representations and toxicity index (TI) model. The isobole analysis indicated additive
interaction between glyphosate and phenol. With the exception of 20:80 of 2,4-dichlorophenol:glyphosate mixture ratio that
was synergistic, other ratios of the mixture were additive. A synergistic (especially 20:80 4-CP:glyphosate mixture) and
additive interaction was observed for some ratios of glyphosate and 4-chlorophenol mixtures. However, the TI of most
isoboles are within the range of 0.5 – 2.0 and are considered additive. Synergistic and additive interaction of formulated
glyphosate with intermediates of 2,4-D was possible against the dehydrogenase activity of Rhizobium species, an important
soil bacterium. The dynamics of the toxic effects thus would depend on the relative amounts of these compounds.
Keywords Roundup®, Herbicides, Dehydrogenase Activity, Phenols, Toxicity
1. Introduction
Increased application of herbicides for improved and
sustained agriculture has resulted in the contamination of
agricultural soils with organic and inorganic pollutants. One
of most commonly used herbicide is glyphosate
(N-phosphonomethylglycine), a post-emergence herbicide.
Glyphosate is desirable due to its effective control of weeds,
rapid inactivation in soil and low mammalian toxicity [1, 2].
However, environmental concern over glyphosate has grown
due to its undesirable side effects on non-target organisms
and persistence in soil and groundwater [3]. Glyphosate
inhibits amino acid synthesis in bacteria and fungi via the
enzyme, 5-enolpyruvyl-shikimate-3-phosphatase synthase in
shikimic acid pathway [4, 5]. However, applications of
glyphosate at high rate have been reported to stimulate
microbial respiration [6, 7].
Glyphosate is often used in mixture with other herbicides
such as 2,4-dichlorophenoxyacetic acid (2,4-D) amines and
esters to widen spectrum, improve herbicide action and save
* Corresponding author:
xrisokey@yahoo.com (C. O. Nweke)
Published online at http://journal.sapub.org/microbiology
Copyright © 2014 Scientific & Academic Publishing. All Rights Reserved
cost [8, 9]. In one of its commercial formulations, Roundup®,
glyphosate is formulated as isopropylamine (IPA) salt of
glyphosate (36% glyphosate a.i.) and a surfactant,
polyoxyethyleneamine (POEA). The increased toxicity of
this glyphosate formulation in comparison to glyphosate is
attributed to the surfactant component [10]. Field application
of glyphosate at recommended rates have been generally
found to be non- toxic to soil microorganisms [11].
Both antagonistic and synergistic effect of 2,4-D on
glyphosate activity have been reported. Glyphosate and
2,4-D mixture has resulted in reduced herbicide activity
against grass compared to glyphosate applied alone [12, 13]
but 2,4-D enhanced the activity of glyphosate against Rape
[12]. Glyphosate applied with 2,4-D increased leafy Spurge
control by 10-fold after 3 months of exposure compared to
glyphosate alone [8]. Combinations of glyphosate with
2,4-D was generally more effective against cutleaf
eveningprimrose than glyphosate applied alone [14].
Synergistiic effect of 2,4-D and glyphosate mixture against
Brazil Pusley has also been reported by Sharma and Singh
[15].
The environmental fate of these herbicides includes
degradation by plants and microorganisms, hydrolysis and
photolysis. A wide range of microorganisms have been