SHORT COMMUNICATION
J For Res (2005) 10:57–60 © The Japanese Forest Society and Springer-Verlag Tokyo 2005
DOI 10.1007/s10310-004-0102-2
Wen Jie Wang · Yuan Gang Zu · Hui Mei Wang
Takashi Hirano · Kentaro Takagi · Kaichiro Sasa
Takayoshi Koike
Effect of collar insertion on soil respiration in a larch forest measured with
a LI-6400 soil CO
2
flux system
Received: January 30, 2004 / Accepted: April 27, 2004
W.J. Wang · Y.G. Zu · H.M. Wang
Key Laboratory of Forest Plant Ecology, Northeast Forestry
University, Harbin, PR China
T. Hirano
Graduate School of Agriculture, Hokkaido University, Sapporo,
Japan
K. Takagi · K. Sasa · T. Koike (*)
Hokkaido University Forests, FSC, Hokkaido University, Sapporo
060-0809, Japan
Tel. +81-11-706-3854; Fax +81-11-706-3450
e-mail: tkoike@exfor.agr.hokudai.ac.jp
Abstract Little information is available on the effect of root
cutting by the collar pre-insertion technique on soil respira-
tion. In this study, we found that soil respiration rates de-
creased with increasing depth of collar insertion in both the
“with live roots intact” and “with live roots severed” treat-
ments, but the rate of decrease was substantially higher in
the former. The cutting of roots, especially fine roots, may
be responsible for this result.
Key words Larch (Larix gmelinii) · Soil respiration rate ·
Collar insertion · Root biomass
Introduction
Soil respiration rate can now be made routinely with a
LI-6400 soil CO
2
flux system (LiCor, Lincoln, NE, USA).
The soil chamber of the system is vented to equilibrate the
pressure inside and outside the chamber, since pressure
differences can have a strong effect on estimates of soil
respiration (Nakayama 1990; Norman et al. 1992; LiCor
1997). Further improvements in soil respiration measure-
ments can be achieved by the insertion of thin-walled
polyvinyl chloride (PVC) collars prior to taking the mea-
surements (Norman et al. 1992). These collars minimize soil
surface disturbance and reduce the sudden flushing of CO
2
in the soil and litter. Despite these improvements, there are
still large variations in soil respiration even within a single
forest, due to biotic and abiotic factors (Lloyd and Taylor
1994; Fang et al. 1998; Davidson et al. 1998; Buchmann
2000; Fang and Moncrieff 2001). One of these factors is the
contribution of root respiration, which can be 40%–60% of
the total soil respiration rate (Bowden et al. 1993; Hanson
et al. 2000). It is, therefore, critical to determine the effect of
inserting the root-cutting collar on soil respiration. How-
ever, available information on the effect of the collars on
soil respiration measurement is still limited (Hanson et al.
2000). Ignoring the effect of the root-cutting collar can
therefore result in misleading soil respiration rates. The
problem is expected to be more serious in forests with a
shallow root system, such as those dominated by Larix
gmelinii. These forests are widespread in central Siberia,
Russia (Kajimoto et al. 2003) and northeast China (Wang et
al. 2002). To examine the effect of inserting the root-cutting
collar on soil respiration, a long-term CO
2
flux site in a L.
gmelinii forest in northeast China was selected as the study
site (Wang et al. 2002). Since soil trenching is a method
commonly used for estimating root respiration (Hanson et
al. 2000; Lee et al. 2003), we also investigated the effect of
collar insertion in such a soil trench.
Study site and methods
Study site
Measurements were made in a larch forest (L. gmelinii
(Rupr.) Rupr.) 36 years old in 2003, located in the Laoshan
Experimental Forest of the Northeast Forestry University
in China (45°20'N, 127°34'E). The mean annual precipita-
tion at Laoshan Experimental Forest was 723 mm and the
annual mean air temperature was 2.8°C. The density of the
larch stand was 1150 stems ha
-1
. The mean height was 18 m
and mean diameter at breast height (DBH) was 17.2 cm.
The soil here is characterized as a dark-brown forest soil
and with about 10% soil organic matter, and pH values of
5.3–6.0. Depth of the litter layer at our study plot was about
2 cm. A more detailed description of this research site is
given by Shi et al. (2001).