Plant Pathology (2002) 51, 794
794 © 2002 BSPP
Blackwell Science, Ltd Oxford, UK PPA Plant Pathology 0032-0862 Plant Pathology, 2002 51 Disease Report Disease Report Disease Report
NEW DISEASE REPORT
First report of the natural occurrence of eight viruses
affecting soybeans in Iran
A. R. Golnaraghi
a
*†, N. Shahraeen
b
, R. Pourrahim
b
, S. Farzadfar
b
and A. Ghasemi
c
a
Research and Science Branch, Islamic Azad University, PO Box 14515-775, Tehran;
b
Plant Virology Department, Plant Pests and Diseases
Research Institute, PO Box 19395-1454, Tehran; and
c
Oilseed Research and Development Co., Tehran, Iran
During the summer of 1999/2000 a survey of viruses
infecting soybeans was conducted in five main soybean
cultivation provinces of Iran (Golestan, Mazandaran,
Ardebil, Lorestan and Khuzestan). A total of 3110 leaf
samples were collected randomly from 80 soybean
fields (30 –50 samples per field). Samples were tested by
the enzyme-linked immunosorbent assay (ELISA) for
the presence of 13 viruses using antisera against Alfalfa
mosaic virus (AlMV), Bean yellow mosaic virus (BYMV),
Cucumber mosaic virus (CMV), Soybean mosaic virus
(SMV), Tobacco mosaic virus (TMV), Tobacco ringspot
virus (TRSV) and Tomato ringspot virus (ToRSV) (Sanofi
Pytho-Diagnostics, 92430 Marnes-la-Coquette, France);
Bean common mosaic virus (BCMV) (Adgen Ltd,
Auchincruive, KA6 5HW, UK); Tobacco streak virus
(TSV) (As-0171, PV-0309) and Tomato spotted wilt virus
(TSWV) (As-0526, As-0580, PV-0526) (DSMZ, BBA
Messeweg 11/12, D-38104 Braunschweig, Germany); Pea
enation mosaic virus-1 (PEMV-1) (Iranian pea isolate
provided by S. Farzadfar); Blackeye cowpea mosaic virus
(BlCMV) (from Dr Pinnow, Department of Plant Pathology
and Physiology, Clemson University, SC 29631, USA);
and Peanut mottle virus (PeMoV) (Seed Pathology Insti-
tute, Flakkejerg, DK-4200 Slagelse, Denmark). The latter
two viruses were tested for by immunodot ELISA, and the
other viruses by double antibody sandwich-ELISA.
Laboratory tests results indicated that the overall virus
disease incidence was 15·7%. SMV was the most com-
mon virus, with an average of 13·3%, followed by TSWV
(5·4%), TRSV (4·2%), TSV (4·1%), PEMV (2·9%),
BYMV (2·2%), ToRSV (2·1%), TMV (1·5%), AlMV
(1·3%), BCMV (0·8%) and CMV (0·6%). In addition,
BlCMV was detected in selected samples with symptoms
from Mazandaran and Lorestan, and PeMoV in samples
from Golesta.
Field symptoms associated with virus infection included
mosaic, mottling, crinkle, vein clearing and vein necrosis
with SMV; mosaic and mottling with AlMV, BCMV,
BlCMV, BYMV, PEMV, PeMoV and TMV; top necrosis
in most TRSV and ToRSV-infected plants; and chlorosis
and necrosis with TSWV. Plants were frequently infected
with more than one virus, however, so it was not possible
to correlate specific symptoms with a particular virus.
SMV, TSV, TRSV, ToRSV and TSWV have previously
been recorded in soybean in Iran (Eskandari, 1978;
Rahimian et al ., 1995; Golnaraghi et al. , 2001), and
AlMV, BCMV, BYMV, CMV, PEMV-1 and PeMoV have
previously been reported from other hosts (Kaiser et al .,
1971; Shahraeen & Bananej, 1995). However, this is the
first report of naturally occurring infections of AlMV,
BCMV, BlCMV, BYMV, CMV, PEMV-1, PeMoV and
TMV in soybean fields in Iran.
References
Eskandari F, 1978. Studies on soybean mosaic in Iran.
Zeitschrift für Pflanzenkrankheiten und Pflanzenschutz 85,
686–8.
Golnaraghi AR, Shahraeen N, Pourrahim R, Ghorbani Sh,
Farzadfar Sh, 2001. First report of Tomato spotted wilt virus
on soybeans in Iran. Plant Disease 85, 1290.
Kaiser WJ, Mossahebi GH, Okhovat M, 1971. Alternate hosts
of viruses affecting food legumes in Iran. Iranian Journal of
Plant Pathology 7, 85.
Rahimian H, Hamdollah-Zadeh A, Montazeri M, 1995. Viruses
associated with soybean pod set failure syndrome in Iran. In:
Proceedings of the 12th Iranian Plant Protection Congress,
1995, Karaj, Iran, 109.
Shahraeen N, Bananej K, 1995. Occurrence of Peanut mottle
virus in Gorgan province. In: Proceedings of the 12th Iranian
Plant Protection Congress, 1995, Karaj, Iran, 110.
*To whom correspondence should be addressed.
†E-mail: agolnaraghi@yahoo.com
Accepted 15 March 2002 at www.bspp.org.uk/ndr where
figures relating to this paper can be viewed.
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