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luteinizing hormone. FASEB J 3:A1474.
16. Weigent DA, Baxter JB, Wear WE, et
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18. Hiestand PC, Mekler P, Nordmann R, et
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Wieconoln Proficiency Program for
Lyme Disease:
A Program for Improving Detection of
Antibodies to Be555rli4 JpurfpJe f ri
Lori L. Bakken, Steven M. Callister, Kay L. Case, and
Ronald F. Schell
Wisconsin State Laboratory of Hygiene, Madison, Wisconsin, and Gundersen Medical
Foundation and Lutheran Hospital, La Crosse, Wisconsin
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1986.
B orrelia burgdorferi, the spirochete
that causes Lyme disease, can
p
infect many nssues of the body
including the skin, heart, nervous system,
and joints.17 Because of this, Lyme dis-
ease symptoms can resemble those of
other illnesses such as influenza, aseptic
meningitis, multiple sclerosis, or rheuma-
toid arthritis. This ability to cause symp-
toms similar to other diseases can make
the accurate diagnosis of Lyme disease
difficult. Therefore, the physician relies
heavily on the ability of the laboratory to
detect accurately antibodies to B. burg-
dorferi.
Serologic testing is currently the only
practical method for establishing a Lyme
disease diagnosis. Infected humans pro-
duce IgM and IgG antibodies that recog-
nize B. burgdorferi antigens, lgM
antibodies generally peak three to six
weeks following infection, while IgG an-
tibodies peak one to three weeks later.
The concentration of IgG antibody may
then remain elevated for weeks to years
following infection. 4' 16
Western immunoblotting is used by
some laboratories to detect antibodies to
individual antigenic components of B,
burgdorferi. However, its use as a prac-
tical clinical laboratory test remains un-
clear. For example, some investigators
have shown that this test is superior to
the indirect ELISA for determining early
Lyme disease, 7 while others have demon-
strated that it is not the method of choice
0197-1859/91/$0.00 + 2.20
for diagnosing early Lyme disease. 2'9
Until these concerns are resolved, this
procedure appears to have limited useful-
ness in most routine diagnostic laborato-
ries.
Currently, indirect immunofluorescent
assays (IFAs) and enzyme-linked immu-
nosorbent assays (ELISAs) are the most
commonly used methods for identifying
and quantifying the presence of antibodies
to B. burgdorferi. Some investigators
have reported that ELISA is more sensi-
tive than IFA, 7 while others report that
IFA is as predictive as ELISA if the labo-
ratory has experience and conservative
cutoff titers are used.12'13 Regardless,
both systems remain somewhat less than
perfect because of both common and
unique problems associated with each.
Whole borrelial cells are invariably
used as the antigen in IFA. However,
this is where similarities among IFA test
procedures end since neither the antigens
nor the methods have been standardized
among laboratories. For example, diag-
nostic titers considered indicative of B.
burgdorferi exposure vary among labora-
tories depending in part on their individ-
ual criterion for intensity of fluorescence.
This causes significant cutoff points to
vary.
Using ELISA, which generally utilizes
either whole cells or sonicated cells as
antigen, sera are considered diagnostically
positive if they elicit absorbances 2.5 to 3
standard deviations above the mean ab-
I
© 1991 Elsevier Science Publishing Co., Inc.
sorbance of a group of healthy controls.
This technique has also not been stan-
dardized among laboratories and wide
variations currently exist.
The cultures of B. burgdorferi used as
antigen may also cause differences among
test procedures. We have observed up to
16-fold differences in reported IgM anti-
body levels of individual patient serum
when performing IFA with diagnostic
slides prepared using B. burgdorferi cul-
tured in separate BSK medium containing
different lots of fraction V bovine serum
albumin. 3 It is easy to envision the same
effect occurring with ELISA.
Another variable affecting serologic
testing may be use of other B. burgdor-
feri isolates as antigen. Most laboratories
and commercial companies use the origi-
nal Shelter Island, NY, USA, isolate B31
(ATCC 35210). However, other isolates
are also used. Greene et al. 6 showed that
immunoblot patterns of sera from dogs
naturally infected with B. burgdorferi
were changed depending on which isolate
of B. burgdorferi was used. We have
also shown that different isolates signifi-
cantly change immunoblot patterns of
sera from humans with Lyme disease. 18
In addition, Schwan et al. 14 demonstrated
that noninfectious B. burgdorferi isolate
B31 (multiple passage) had a reduced
immunoblot compared to an infectious
isolate. These observations provide evi-
dence that a serum may be falsely labeled
negative if reactivity varies depending on
I