ICANCER RESEARCH56. 5205-5210, November 15. 9961
ABSTRACT
A marked effect ofcharge modification on the uptake and phototoxicity
of a photoimmunoconjugate (PlC) was demonstrated. A site-specific con
jugation strategy was developed to attach the photosensitizer chlorin e6
(ce6)to the F(ab')2 fragment of the murine antiovarian cancer monoclonal
antibody 0C125. Poly-L-lysine linkers carrying ce6with a cationic charge
or by polysuccinylation with an anionic charge were used and covalently
attached to partially reduced antibody via a heterobifunctional reagent.
PICs were purified by column chromatography and were also radiola
beled with 1251.PlC binding and uptake were studied with a human
ovarian cancer cell line, NIH-OVCAR-5, and a nonantigen-expressing
colon cancer cell line, SW1116, and the data were compared with the
binding and uptake of nonspecific rabbit IgG PICs. PICs with both
cationic and anionic charges preserved antigen binding as shown by
competition studies with native antibody, but the cationic PlC had up to
17 times higher cellular uptake of ceo, probably due to enhanced inter
nalization. The ratio of ce, to ‘@I retained by the cells varied with the
likelihood ofinternalization and lysosomal degradation. The phototoxicity
ofthe PICs generally varied with their uptake, but a correlation was found
between lysosomal hydrolysis as measured by an increased cellular ratio
ofce,:'251 and increased relative phototoxicity. These data suggest cationic
PICs may have advantages for photoimmunotherapy of disseminated
intracavity cancer following local administration.
INTRODUCTION
Mabs3 directed against tumor-associated antigens are used to target
drugs or radioisotopes to tumors for both diagnosis and therapy (1).
Photoimmunotherapy, in which PSs are conjugated to Mabs has
recently been explored (2—5).These PSs produce cytotoxic species
(singlet oxygen or radicals) when irradiated with red light. This
approach using PICs has inherent dual selectivity: in addition to the
tumor-targeting properties of the Mab, tissue destruction is restricted
to the field of illumination. This modality may be suitable for treat
ment of small diffuse malignancies present in a cavity such as the
peritoneum or bladder (6). Therapeutic success using Mab conjugates
has been modest due to (a) limited uptake and internalization of
conjugates and (b) lack of specificity. It is well known that the
introduction of positive charges onto macromolecules increases the
rate of endocytosis (7, 8), and we hypothesized that this would apply
to PICs and that a positively charged PlC would have greater tumor
uptake when administered intracavitary. Uptake of the cationized
PICs could be further enhanced by the use of F(ab')2 fragments, which
have the advantages of greater penetration into the tumor and faster
clearance (9).
This study tests the above hypothesis and is a first report on (a) the
synthesis of site-specific PICs using a strategy which is equally
Received 5/8/96; accepted 9/16/96.
The costs of publication of this article were defrayed in part by the payment of page
charges. This article must therefore be hereby marked advertisement in accordance with
18 U.S.C. Section 1734 solely to indicate this fact.
I This work was supported by NIH Grant RO1 AR40352 and in part by the Department
of Defense Free Electron Laser Program facilities.
2 To whom requests for reprints should be addressed. Phone: (617) 726-6996; Fax:
(617) 726-3192.
3 The abbreviations used are: Mab, monoclonal antibody; PS, photosensitizer; PlC.
photoimmunoconjugate; NHS, N-hydroxysuccinimide; ce,, chlorin ,.,,; p1, poly-L-lysine;
succ, succuccinylated; SPDP, 2-pyridyl-dithiopropionic acid amide.
applicable to intact IgG or to F(ab')2 fragments and which has the
option of preparing PICs bearing either cationic or anionic charges
and (b) the effect of charge modification of PICs constructed from the
F(ab')2 fragment of the Mab OCl25 on their interaction with a human
ovarian cancer cell line, NIH:OVCAR-5, expressing the target antigen
CA125 (10).
MATERIALS AND METHODS
Cell Line and Mab. NIH:OVCAR-5(OVCAR-5) cells were purchased
from Dr. T. Hamilton (Fox Chase Cancer Institute, Philadelphia, PA) and
SW1116 human colon cancer cells were purchased from American Type
Culture Collection (Rockville, MD). Cells were grown in RPM! 1640 contain
ing HEPES, glutamine, 10% heat-inactivated fetal bovine serum, 100 units/ml
penicillin, and 100 pg/mI streptomycin and maintained in an incubator at 37°C
in an atmosphere of 5% carbon dioxide. OC125F(ab')2 murine Mab was a
generous gift from Centocor (Malvern, PA). Rabbit lgG was obtained from
Sigma (St. Louis, MO).
Radiolabeling. OC125F(ab')2 and rabbit IgG were radiolabeled with 1251
using the lodo-Gen procedure (I 1). ‘251-labeled OC125F(ab')2 had a specific
activity of 8.7 MBq/mg whereas ‘251-labeled rabbit IgG had a specific activity
of2.l MBq/mg.
Preparation of Poly-L-lysine Ce6 Conjugates. All reactions were carried
out in the dark at room temperature. The NHS ester of ceo was prepared by
reacting 1.5 equivalents of dicyclohexylcarbodiimide and 1.5 equivalents of
NHS with 1 equivalent of ceo in dry DMSO for 24 h and was frozen in aliquots
for further use. p1 hydrobromide (average Mr 25,000 50 mg) was dissolved in
dry DMSO (50 ml) and N-ethylmorpholine (1 ml) was added followed by
ceo-NHS ester (25 mg) and reacted for 24 h. The solution of pl-c@6 obtained
was placed into a DMSO-resistant dialysis membrane of Mr 3500 cutoff
(Spectrum Medical Industries, Los Angeles, CA) and dialyzed twice against 5
liters of I mM potassium phosphate buffer (pH 9).
The pl-c,@ dissolved in DMSO was treated with an excess of succinic
anhydride (100 mg dissolved in 0.5 ml of dry DMSO, 4-fold excess for each
primary amino group) for 24 h to afford the anionic pl-ceo-succ which was
dialyzed as above.
Preparation of Site-specific PICs with OC12SF(ab')2 and Rabbit IgG.
The DMSO solution of p1-c@6 was treated with a solution of pyridyldithiopro
pionic acid-NHS ester in DMSO (3 equivalents/pI chain). After reacting for
24 h, it was dialyzed twice against 5 liters of 1 mist acetic acid. OC125F(ab')2
fragment or rabbit IgG (5 mg) was dissolved in S ml of 10 miss phosphate
buffer (pH 7.5) containing I msi EDTA and 5 mM mercaptoethylamine
hydrochloride, and the solution was purged with nitrogen and shaken in the
dark for 2 h. The solution was then dialyzed three times against 5 liters of 10
mM phosphate buffer (pH 7.5); 1 msi EDTA. To this solution was added 3
equivalents of pl-ceo-SPDP and left 24 h before purification.
To the pl-cen-SPDPsolution described above was added a 4-fold excess of
succinic anhydride. After 24 h the resulting p1-c@6-SPDP-succ was dialyzed
(three times) against 5 liters of 1 mMsodium acetate (pH 9) and then added to
the partially reduced Mab as described above.
PlCs were purified on Sephadex G200 columns and characterized by
SDS-PAGE (12). Quantities of PICs were calculated by measuring absorption
spectra in 0.1 MNaOHI1% SDS solution and assuming the € (400 nm) of c,.,,,
1.5 X l0@,remained unchanged by conjugation.
Fluorescence Localization in Living Cells. OVCAR-5 cells were grown
(60% confluent) in P35 dishes containing 4 ml of medium and a 22-mm square
glass coverslip. PICs were added at a final concentration of 2 pM cen equivalent
in serum-containing medium and allowed to incubate for 3 h. The coverslips
were washed three times with PBS and put on histological slides in PBS and
examined with an epiillumination fluorescence microscope (Axiophot; ZeiSS,
5205
Effect of Charge on the Interaction of Site-specific Photoimmunoconjugates with
Human Ovarian Cancer Cells'
Michael R. Hamblin, Jaimie L. Miller, and Tayyaba Hasan2
Wellman Laboratories of Photomedicine, Department of Dermatology, Massachusetts Genera! Hospital. WEL224, Harvard Medical School. Boston, Massachusetts 02114
on July 13, 2017. © 1996 American Association for Cancer Research. cancerres.aacrjournals.org Downloaded from