ELSEVIER Electroencephalography and clinical Neurophysiology 101 (1996) 32-41
Abnormal excitability of the corticospinal pathway in patients with
amyotrophic lateral sclerosis: a single motor unit study using transcranial
magnetic stimulation
N. Kohara a,*, R. Kaji a, y. Kojima a, K.R. Mills c, H. Fujii a, T. Hamano a, j. Kimura a,
N. Takamatsu b, T. Uchiyama b
a Departmentof Neurology, Kyoto UniversityHospital, Shogoin, Sakyo-Ku, Kyoto 606, Japan
b NEC-Sanei, Kyoto, Japan
¢ UniversityDepartment of Clinical Neurology, The Radcliffe Infirmary, Oxford, UK
Accepted for publication: 19 July 1995
Abstract
The pathophysiology of corticospinal tract degeneration in amyotrophic lateral sclerosis (ALS) was investigated by studying the effect
of transcranial magnetic stimulation on discharge characteristics of single motor units during voluntary activation. The motor units were
recorded from the first dorsal interosseus muscles of 12 patients with ALS, 14 healthy subjects, 12 patients with upper motor neuron
lesions and 9 with pure lower motor neuron diseases. More than 100 magnetic stimuli were delivered over the scalp during minimal
muscle contraction. The occurrence of motor unit discharges was plotted in a peristimulus time histogram. An increase in discharge
probability at latencies of 20-30 msec, that represents monosynaptic activation (primary peak) was found in normal units. Motor units
from ALS patients with short disease durations had significantly increased discharge probabilities in the primary peak (P < 0.01). Motor
units from 4 ALS patients with upper motor neuron signs showed double primary peaks: an initial synchronized peak followed by a
dispersed peak. The latter was ascribed to a slow corticospinal pathway, which remains undetected or is functionally insignificant in
healthy subjects. We conclude that the excitabilities of the surviving corticospinal tract pathways are abnormally increased in ALS,
especially in the early stage.
Keywords: Peristimulus time histogram (PSTH); Amyotrophic lateral sclerosis (ALS); Single motor unit; Excitability; Corticospinal tract
1. Introduction
Amyotrophic lateral sclerosis (ALS) is a fatal disease of
unknown etiology. It involves both the upper and lower
motoneurons, of which those comprising the direct corti-
cospinal system seem to be selectively involved (Hudson
and Kiernan, 1988; Eisen et al., 1992; Kaji and Kimura,
1993).
Mounting clinical, pathological and physiological evi-
dence points to the progressive loss of motoneurons, but
little is known about the interaction of the upper and lower
motoneurons in the disease process. The advent of trans-
cranial magnetic or electrical stimulation of the motor
* Corresponding author. Tel.: + 81 75 7513772; Fax: + 81 75 7619780.
cortex has made possible functional assessments of the
corticospinal tract system. Previous studies that used sur-
face recordings of compound muscle action potentials after
cortical stimulation have shown prolonged conduction
times from the cortex to the spinal level and decreased
amplitude or complete lack of evoked muscle responses in
ALS (Ingram and Swash, 1987; Schriefer et al., 1989;
Eisen et al., 1990; Berardelli et al., 1991). These findings
are not, however, specific to ALS. They have been re-
ported in such diseases that affect upper motoneurons as
multiple sclerosis (MS) (Hess et al., 1987; Berardelli et al.,
1988; Ingram et al., 1988) and cerebrovascular disease
(CVD) (Bridgers, 1990; Berardelli et al,, 1991).
Characteristic findings of ALS, such as fasciculations
and muscle cramps, sometimes antedate development of
muscle weakness and atrophy (Tyler and Shefner, 1991).
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