Decomposition of a protein solution into Voronoi shells and Delaunay layers
Kim A.V
1
, Voloshin V.P.
1
, Medvedev N.N.
1,2
1
Institute of Chemical Kinetics and Combustion, SB RAS,
630090 Novosibirsk, Russia
2
Novosibirsk State University, Novosibirsk, Russia
e-mail: nikmed@kinetics.nsc.ru
Geiger A.
3
3
Physikalische Chemie, Technische Universität
Dortmund, 44221 Dortmund, Germany
e-mail: alfons.geiger@udo.edu
Abstract— A simple formalism is proposed for a quantitative
analysis of interatomic voids inside and outside of a molecule
in solution. It can be applied for the interpretation of
volumetric data, obtained in studies of protein folding in water.
The method is based on the Voronoi-Delaunay tessellation of
molecular-dynamic models of solutions. It is suggested to select
successive Voronoi shells, starting from the interface between
the solute molecule and the solvent, and continuing to the
outside (into the solvent) as well as into the inner of the
molecule. Similarly, successive Delaunay layers, consisting of
Delaunay simplexes, can also be calculated. Geometrical
properties of the selected shells and layers are discussed. The
behavior of inner and outer voids is discussed by the example
of a molecular-dynamic model of an aqueous solution of the
polypeptide hIAPP.
Index terms—Voronoi diagram, solvation shell, molecular
dynamics of solutions, Voronoi cells, Delaunay simplexes.
I. INTRODUCTION
The investigation of biological molecules in aqueous
solution is an important problem of molecular biology. In
particular, it is important to understand the mechanism of
protein folding in water at different temperatures and
pressures. Volumetric experiments are used for this study.
The change of the volume of the solution, induced by the
addition of a protein molecule is measured [1]. The influence
of temperature and pressure induces changes of the
volumetric properties, both inside the solute molecule, at its
boundary, and also in the surrounding water. The knowledge
of these contributions to the volume of the solution helps to
validate propositions about the occurring conformational
changes. However, using only experimental data, it is very
difficult to separate these contributions. Computer
simulations help to solve this problem.
Models of the solutions are generated usually by
molecular dynamic simulations, see for example [2]. The
next step is the analysis of the models: detection and
characterization of interatomic voids and local densities.
There are very different approaches used for the analysis of
voids in atomic and molecular systems. Some of them were
developed for the investigation of the empty space between
the atoms in liquids and glasses [3-5], granular matters and
colloids [6,7], polymers and membranes [8,9]. Others are
specialized to study cavities and pockets in large biological
molecules [10-12]. Solvation shells [13,14] and the boundary
region between proteins are also studied [15,16].
Consecutive shells, consisting of Voronoi cells, were used
for the analysis of the density of hydration shells around
polypeptides in [17]. However, we are not aware of articles,
where the voids both inside and in the surroundings of a
solute molecule were analyzed. Such investigations should
be made by a single-stage method for all regions of the
solution. Fortunately, there is no necessity to develop a new
method for such a work. At present, there is no doubt, that
the most suitable and general method for the selection and
analysis of voids and the local density in molecular system is
an approach, which is based on Voronoi diagrams (the
Voronoi -Delaunay method) [18,19].
In this work, we present a simple technique for the
decomposition of the Voronoi-Delaunay tessellation into
shells (layers) related with the solute. It allows to
characterize voids (local density) both inside and outside the
solute molecule.
II. VORONOI-DELAUNAY TESSELLATION OF A SOLUTION
Fig.1 shows a two-dimensional illustration of a solution
model and its Voronoi-Delaunay tessellation.
Remember, the size of the atoms should be taken into
account, if one studies interatomic voids [3,20,21]. This
means that the Voronoi tessellation should be calculated
allowing for the surface of the atoms. Thus we should deal
with S-tessellation [22,23] (additively weighted [18]),
instead of the ordinary Voronoi tessellation (related with the
atomic centers). In this case we make a correct assignment of
the empty space to a given atom, i.e. we include all points of
space, which are closer to the surface of a given atom, than
to the surfaces of all other atoms of the system. A simpler
variant, which considers the atomic surfaces, is the well-
known power or radical tessellation [18,21,24]. In this case
the assignment of the empty space to individual atoms is not
quite physical, but it is easier to implement. The known
complexities of the S-tessellation (theoretically possible
disconnectivity of the tessellation and overlapping of
Delaunay simplexes in some cases [19,23,25] do not arise for
our molecular systems, where the size difference of the
atoms is rather small (usually less than a factor of 2). In
addition, these peculiarities of the S-tessellation can be easily
taken in to account at the calculation of the tessellation.
The molecules of the solvent (usually water molecules)
are considered as uniform spheres, as it is usually done in
structure analyses of computer models of water and water
solutions. Note, the specific features of the interaction
2012 Ninth International Symposium on Voronoi Diagrams in Science and Engineering
978-0-7695-4724-4/12 $26.00 © 2012 IEEE
DOI 10.1109/ISVD.2012.18
95