Journal of Aquatic Science, 2015, Vol. 3, No. 1, 14-18
Available online at http://pubs.sciepub.com/jas/3/1/3
© Science and Education Publishing
DOI:10.12691/jas-3-1-3
The Daily Light-dark Cycle of Photosynthetic Oxygen
Evolution in Three Species of Tropical Calcareous Algae
Alvin P. Monotilla
*
, Serafin M. Geson, Paul Isaac O. Dizon, Danilo T. Dy
Marine Biology Section, University of San Carlos, Cebu City, Philippines
*Corresponding author: alvinmonotilla@yahoo.co.jp
Abstract Endogenous coordination between light, temperature and other factors by different species of algae
would be vital in the production of several proteins needed for growth and adaptations, and therefore will affect their
productivity. Among plant activities that are governed by daily light-dark cycles (i.e. 12L:12D and 12L:12L) among
calcareous algae (Halimeda simulans, Mastophora rosea and Padina australis) were conducted by monitoring the
dissolved oxygen (DO) concentration inside incubating bottles with algal samples for 24h and the features of the
resulting DO curves were determined by measuring the number of pixel under the curve through image analysis.
There was no significant difference among algae during the initial 12L:12D cycle suggesting their normal response
to light-dark cycles. However, after six days under continuous light, M. rosea showed a significant decrease in the
DO curve (lesser number of pixel under the curve) compared to DO curves during the initial 12L:12D cycle. The
decrease in the DO concentration during the continuous L:L treatments might be attributed to the photoinhibitory
effect of the red alga being less adoptive to subsequent high intensities. Although an increase in DO concentrations
is expected with continuous light, not all algae responded to it. Only Padina exhibited circadian rhythm in our 24h
observation under continuous light.
Keywords: Coral reef, Padina, Halimeda, Mastophora, circadian rhythm
Cite This Article: Alvin P. Monotilla, Serafin M. Geson, Paul Isaac O. Dizon, and Danilo T. Dy, “The Daily
Light-dark Cycle of Photosynthetic Oxygen Evolution in Three Species of Tropical Calcareous Algae.” Journal
of Aquatic Science, vol. 3, no. 1 (2015): 14-18. doi: 10.12691/jas-3-1-3.
1. Introduction
There are behaviors and processes in organisms that are
observed to be repetitive and therefore predictable during
the course of their existence. The adaptation of living cells
to daily fluctuations of light, over a period of about a day,
is reflected by endogenous rhythms (circa diem) [3].
These rhythms are thought to be regulated by internal
oscillators (circadian clocks) or are endogenous biological
clock that governs biochemical phenomenon or behavior
which allows organisms to predict environmental changes,
such as day/night and seasons that result from the rotation
of the earth [19]. Among the many environmental cues,
light affects virtually all aspects of plant growth and
development like gravitropism, phototropism, floral
initiation, germination and vegetative circadian rhythm to
daily changes in environmental conditions over a 24h light
dark (L:D) cycles (Kendrick and Kroneberg, 1994 in [2]).
It is for this reason that studies on the effects of light on
circadian rhythm and daily L:D cycles in plant are needed
to maximize their productivity potential or their
sustainability toother geographic locations.
Similar to land plants, circadian rhythms are also
expressed in marine algae under various light regimes and
irradiances. The role of circadian rhythm has been very
evident on the high productivity of some algae in the
initiation and formation of proteins vital in the synthesis
of organic compounds needed for growth and other
functions. Various activities tested in rhodophytes
Kappaphycus alvarezii [10], Gracilaria tenuistipitata [14]
and Gracilaria chilensis [5] on nitrate assimilation
characteristics revealed that nitrate reductase activity is
under circadian control, exhibiting daily rhythm under L:D
cycles and constant light. On the other hand, Drew and Abel
[7] observed that the segments of Halimeda rapidly become
very pale after dark, but are green enough by dawn to
permit rapid photosynthesis as soon as light is available
suggesting that the re-emergence of the chloroplast before
dawn and their subsequent withdrawal appears to be
controlled by an endogenous rhythm which is independent
of light and reportedly work at temperatures ranging
between 20°C and 30°C [9]. Photosynthesis and other
processes (i.e., calcification in calcareous algae) appear to
be under circadian control in some of the algae studied.
Larkum et al., [12] concluded that in Halimeda macroloba,
alkalization of the internal utricular space triggers
calcification during photosynthesis and exhibited an
endogenous rhythm during a 12:12 hrs L:D treatment but
not in continuous light. Unlike Halimeda, another green
alga, Acetabularia exhibited endogenous rhythm even in
continuous light [11]. This contradiction indicates that
further studies are needed to verify the effects of
continuous light on other calcareous or coralline algae.
The polarographic dissolved oxygen (DO) meter can be
used to monitor oxygen evolution and consumption of an
incubated algal sample. With appropriate methodology