Open Agriculture. 2018; 3: 25–31
Keywords: Gene effects; Heritability; Heterosis; Generation
means analysis
1 Introduction
Triticale (x Triticosecale Wittmack) in Argentina is
potentially suitable to various uses, although the most
frequent is direct grazing, it can also be used as a dual
purpose and to supplement or include in rations. Triticale
is in a free expansion period, especially in sub-humid and
semi-arid areas where it successfully replaces the use of
rye (Grassi et al. 2004). Its rusticity is similar to rye but
with superior forage quality (Amigone and Kloster 1997).
The selection of experimental line for grain yield registers
varied antecedents in Argentina (Grassi et al. 2003, 2004;
Cardozo et al. 2005; Castro et al. 2011; Ferreira et al. 2015).
Grain yield is a complex trait with multiple
components and is strongly influenced by the
environment, in which numerous morphological and
physiological traits contribute in its expression. It would
be necessary to know the inheritable and not inherited
contribution with the help of genetic parameters (Singh
and Chaudary 1985; Kahrizi et al. 2010; Maniee et al.
2009). The analysis of the nature and magnitude of the
gene effects on quantitative traits helps the researcher to
design an efficient improvement program and to provide
information on the potential manifestation of heterosis
(Shekhawat et al. 2000). Analysis of relative mid-parent
heterosis (Mh%) and heterobeltiosis (Hh%) allows to
define the breeding strategy (Yildirim et al. 2014). Recent
studies in triticale suggest that hybrids performance are
significantly higher than cultivated varieties (Oettler et al.
2003; 2005; Yildirim et al. 2014).
Generation mean analysis is a simple technique to
estimate the genetic effects and to elucidate the nature
of the epistasis, being useful to decide the procedure to
be adopted in the improvement of quantitative traits
(Abd El-Rahman and Hammad 2009). Joint-scaling test
https://doi.org/10.1515/opag-2018-0003
received October 9, 2017; accepted December 14, 2017
Abstract: Triticale (x Triticosecale Wittmack) is a cereal
used in Argentina as a winter seasonal pasture or feeding
grain. Efforts made on breeding have led to varieties
with potential grain yield similar or greater than with
wheat. The aim of this study was to determine the gene
effects, heritability, genetic gain, heterosis and inbreeding
depression in various quantitative traits of a cross from
contrasting lines of triticale using generation means
analysis. The thousand kernel weight was adjusted to a
model of additive-dominance, while the rest of the traits
exhibited significant epistatic effects: additive x additive
(i) for hectoliter weight, additive x dominance (j) for
number grains per spike and grain yield per plant. Other
traits showed a more complex as inheritance additive
x additive (i) and additive x dominance (j) interactions
effects for spike length or additive x dominance (j) and
dominance x dominance (l) interactions effects for
number of spikes per plant and number of spikelets per
spike. High values of heritability and genetic gain were
also detected for grain yield, spikelets per spike, spike
per plant and hectoliter weight. All traits except spike
per plant had positive relative mid-parent heterosis, but
none resulted in positive heterobeltiosis, therefore this
cross is not recommended for obtaining hybrids varieties.
Selection in early generations by thousand kernel weight
and hectoliter weight would be efficient, while the
selection in advanced generations where segregation is
minimal would be recommended for the remaining traits.
Research Article
Enzo David Ferrari*, Víctor Aníbal Ferreira, Ezequiel Martín Grassi,
Aurora María Teresita Picca, Héctor Antonio Paccapelo
Genetic parameters estimation in quantitative
traits of a cross of triticale (x Triticosecale W.)
*Corresponding author: Enzo David Ferrari, Faculty of Agronomy,
National University of La Pampa, Santa Rosa, La Pampa, Argentina,
E-mail: ferrarienzo@agro.unlpam.edu.ar
Aurora María Teresita Picca, Héctor Antonio Paccapelo, Faculty of
Agronomy, National University of La Pampa, Santa Rosa, La Pampa,
Argentina
Víctor Aníbal Ferreira, Ezequiel Martín Grassi, Faculty of Agronomy
and Veterinary, National University of Río Cuarto, Río Cuarto, Córdo-
ba, Argentina
Open Access. © 2018 Enzo David Ferrari et al., published by De Gruyter Open. This work is licensed under the Creative Commons Attribution-
NonCommercial-NoDerivs 4.0 License.
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