IEEE JOURNAL OF SELECTED TOPICS IN QUANTUM ELECTRONICS, VOL. 22, NO. 3, MAY/JUNE 2016 6801706
Functional Connectivity During Phonemic and
Semantic Verbal Fluency Test: A Multichannel
Near Infrared Spectroscopy Study
Chun-Jung Huang, Po-Han Chou, Hao-Lin Wei, and Chia-Wei Sun
Abstract—Verbal fluency tests (VFTs) are widely used frontal
lobe neuropsychological tests. They have been frequently used in
various functional brain mapping studies. There are two versions
of VFTs based on the type of cue: the letter fluency task (LFT) and
the category fluency task (CFT). However, the fundamental aspects
of brain connectivity across the frontotemporal regions during the
VFTs have not been elucidated. In this study, we hypothesized
that differences in cortical functional connectivity over the left and
right frontotemporal regions may be observed by means of multi-
channel functional near-infrared spectroscopy (fNIRS) during the
performance of LFT and CFT. Our results from fNIRS (ETG-4000)
showed different patterns of brain functional connectivity during
the two types of VFTs, which was consistent with the different cog-
nitive requirements of each task. We demonstrate increased brain
functional connectivity over the frontal and temporal regions dur-
ing the LFT than during the CFT; these results are in line with
previous brain activity studies using fNIRS to demonstrate higher
frontal and temporal region activation during LFT and CFT, with
more pronounced frontal activation by the LFT.
Index Terms—Verbal fluency tests, functional connectivity of
brain, fNIRS.
I. INTRODUCTION
F
UNCTIONAL near-infrared spectroscopy (fNIRS) is a
recently developed technology for monitoring functional
brain activity based on the absorption and scattering properties
of near-infrared light. fNIRS allows non-invasive measurements
of the spatiotemporal characteristics of neural activity in the
frontotemporal region by measuring changes in the concentra-
tions of oxy-hemoglobin (ΔHbO
2
), deoxy-hemoglobin (ΔHb),
and total hemoglobin (ΔHbT) [1], [2]. fNIRS has several advan-
tages over existing imaging techniques, such as PET, SPECT,
and fMRI, because it is noninvasive, is easy to administer,
Manuscript received July 31, 2015; revised September 21, 2015, September
28, 2015, October 12, 2015, and October 24, 2015; accepted November 11, 2015.
Date of publication December 22, 2015; date of current version April 1, 2016.
This work was supported in part by the Taiwan National Science Council un-
der Grant NSC 101-2628-E-009-026-MY3, Grant NSC 102-2321-B-009-002,
Grant NSC 102-2622-E-009-007-CC3, Grant NSC 102-2627-E-010-001, Grant
NSC 102-3011-P-010-003, and a grant from Ministry of Education, Aim for
the Top University Plan in National Chiao Tung University 101W9866, and
in part by grants from Taichung Veterans General Hospital, Taiwan (TCVGH-
1034001A, TCVGH-1044002B, and TCVGH-YM1040101). Chun-Jung Huang
and Po-Han Chou have contributed equallly to this paper. (Corresponding au-
thor: Chia-Wei Sun.)
C.-J. Huang and C.-W. Sun are with the Department of Photonics, Na-
tional Chiao Tung University, Hsinchu 300, Taiwan (e-mail: mortis1115.eo00@
g2.nctu.edu.tw; chiaweisun@nctu.edu.tw).
P.-H. Chou and H.-L. Wei are with the Department of Psychiatry, Taichung
Veterans General Hospital, Taichung 407, Taiwan (e-mail: phchou1980@gmail.
com; yrbovlxh@yahoo.com.tw).
Color versions of one or more of the figures in this paper are available online
at http://ieeexplore.ieee.org.
Digital Object Identifier 10.1109/JSTQE.2015.2503318
tolerates small movements, is inexpensive, and provides excel-
lent time resolution and moderate spatial resolution [3]. To date,
fNIRS has been applied in many studies to explore the functional
integration among brain regions during different states, includ-
ing resting and task states in normal people and patients with
psychiatric disorders [4]–[9].
Verbal fluency tests (VFTs) are widely used frontal lobe neu-
ropsychological tests and have been frequently used in various
functional brain mapping studies [10]–[13]. There are two ver-
sions of VFTs based on the type of cue used: category fluency
task (CFT) requires the participant to generate words belonging
to a specific semantic category, while during the letter fluency
task (LFT), subjects were asked to retrieve words with certain
phonemic properties, e.g., words beginning with a given let-
ter [14]. These different search strategies pose the question of
whether semantic and phonemic systems for word generation
depend on different neural substrates. Previous studies have
shown that lesions in the temporal lobe may lead to deficits
in semantic function, whereas phonemic function depends on
frontal regions, as well as temporal lobes [15].
There have been several studies investigating the differences
in brain activity during both versions of VFTs in healthy indi-
viduals or patients with schizophrenia [16]–[20]. However, the
fundamental aspects of brain connectivity across spatial regions
of the frontotemporal regions during either versions of the VFT
have not been elucidated to date. To the best of our knowledge,
only one NIRS study investigated functional connectivity during
a VFT, but evaluated a limited area of the cortex [21].
Therefore, the objective of this study was to investigate the
functional connectivity during two versions of the VFTs over a
wider area of the bilateral frontotemporal regions using a 52-
channel continuous wave (CW)-NIRS system (ETG-4000). The
NIRS imaging technique is so-called functional optical topog-
raphy. We hypothesized that differences in cortical functional
connectivity over the left and right frontotemporal regions could
be observed by means of multi-channel fNIRS during LFT and
CFT. Moreover, we expected LFT to cause greater frontal cortex
involvement than CFT due to the increased selection demands
associated with retrieving words based on spelling rules, relative
to word retrieval based on semantic categories.
II. MATERIAL AND METHODS
A. Study Participants
Thirteen healthy individuals (six men and seven women)
without any psychiatric disease screened by Mini-International
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