Molecular Characterization of Enterotoxigenic Escherichia coli: Effect on
Intestinal Nitric Oxide in Diarrheal Disease
Muhammad Arfat Yameen
1
, Ebuka Elijah David
2*
, Humphrey Chukwuemeka Nzelibe
3
, Muhammad Nasir Shuaibu
3
, Rabiu Abdussalam Magaji
4
, Amakaeze
Jude Odugu
5
and Ogamdi Sunday Onwe
6
1
Department of Pharmacy, COMSATS Institute of Information Technology, Abbottabad, Pakistan
2
Department of Biochemistry, Federal University, Ndufu-Alike, Ikwo, Ebonyi State, Nigeria
3
Department of Biochemistry, Ahmadu Bello University, Zaria, Kaduna State, Nigeria
4
Department of Human Physiology, Ahmadu Bello University, Zaria, Kaduna State, Nigeria
5
Medical Laboratory, Ahmadu Bello University, Teaching Hospital, Zaria, Kaduna State, Nigeria
6
Laboratory Service Unit, Federal Teaching Hospital, Abakiliki, Ebonyi State, Nigeria
*
Corresponding author: Ebuka Elijah David, Department of Biochemistry, Federal University, Ndufu-Alike, Ikwo, Ebonyi State, Nigeria, Tel: +2348033188823; E-mail:
david.ebuka@funai.edu.ng
Received date: May 01, 2018; Accepted date: May 25, 2018; Published date: May 30, 2018
Copyright: ©2018 Yameen MA, et al. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted
use, distribution, and reproduction in any medium, provided the original author and source are credited.
Abstract
This study was aimed to investigate the effect of enterotoxigenic E. coli (ETEC)-induced diarrhea on fecal nitric
oxide (NO) and intestinal inducible nitric oxide synthase (iNOS) expression in rats. E. coli isolates were gotten from
infants diarrheal samples. Multiplex polymerase chain reaction (PCR) was used to detect the presence of eltA gene
for heat-stable (ST) enterotoxigenic E. coli and eltB for heat-labile (LT) enterotoxigenic E. coli. Disk diffusion method
was used to determine the antibiotic susceptibility of the organisms. Biofilm formation was detected by thiazoylblue
tetrazolium bromide dye in a 96-well plate. Fecal NO was measured using standard griess reaction system. Reverse
transcription PCR was used to investigate the expression of iNOS expression. Although none of the ETECs isolated
in this study belonged to the classic serotype, serogroup O6 and O8 were found to be associated with ETECs.
Among the three ETECs, two were found to be multidrug resistant. The biofilm production abilities of all the ETECs
were found to exist between weak and moderate biofilm producers. Fecal NO was found to be elevated in both LT
and ST-induced diarrheal groups but there was no corresponding intestinal iNOS expression. This suggests that the
elevated NO could be as a result of up-regulation of constitutive NOS rather than iNOS.
Keywords: Enterotoxigenic E. coli; Escherichia coli; Nitric oxide;
Diarrheal disease
Introduction
Diarrhea is the second leading cause of death in children under the
age of ive of which about 1.5 million die each year. his number
represents nearly one in ive child deaths. More of these children die of
diarrhea than AIDS, malaria and measles combined and greater
percentage of this death due to diarrhea occur in Africa and South
Asia of which Nigeria ranks second, with over 151,700 deaths per year
[1]. Diarrhea can be deined as having watery stools for at least three
times in a day or when it is more frequent than normal in an
individual. It is a symptom of gastrointestinal infections that can be
caused by bacteria, viruses, protozoa and other pathogens. However,
the major organisms that cause most cases of childhood acute diarrhea
are Rotavirus, E. coli, Shigella, Salmonella, Campylobacteria and Vibro
cholera (during epidemics). hese pathogens are transmitted normally
from the stool of one person to the mouth of another through diferent
media like food or water [1]. Rotavirus and pathogenic E. coli are the
most common cause of childhood diarrhea while Shigella spp. remains
the most important causes of acute bloody diarrhea ( dysentery),
accounting for about 15% of death in children under 5 years [2].
Escherichia coli are Gram-negative, oxidase-negative, rod-shaped
bacterium from the family Enterobacteriaceae [3]. hey are
commensal bacteria that can be found in intestinal micro lora of a
variety of animals including man. Not all the strains of E. coli are
harmless since some can cause debilitating and sometimes fatal
diseases in humans as well as mammals and birds [4]. Pathogenic
strains are divided into intestinal pathogens (InPEC) causing diarrhea
and extraintestinal E. coli (ExPEC) causing a variety of infections
including urinary tract infections (UTI), meningitis and septicemia
[5]. InPEC strains of E. coli also known as diarrheagenic E. coli (DEC)
is a major etiological agent of pediatric diarrhea, accounting for over 2
million deaths annually [6]. It continues to be the most common cause
of infantile morbidity and mortality most especially in developing
countries and sub-Saharan Africa [7] particularly in children under 5
years of age [8]. Nigeria records over 50,000 diarrhea-speciic mortality
among children less than 5 years [3]. DEC can be transmitted via the
oral-fecal route by ingesting food or water contaminated by human or
animal feces [7]. An altered movement of ions and water in
gastrointestinal tract is at the heart of diarrheal diseases. InPEC can
alter the balance between luid-electrolyte absorption and secretion
leading to diarrhea [9]. DEC are divided into enteropathogenic E. coli
(EPEC), enterotoxigenic E. coli (ETEC), enterohemorrhagic E. coli
(EHEC), enteroinvasive E. coli (EIEC), enteroaggregative E. coli
(EAEC) and difusely adherent E. coli (DAEC) based on their speciic
virulence factors and phenotypic traits. Each pathological type has
characteristic virulence determinants that contribute to its pathogenic
mechanisms [10]. he prevalence and other epidemiological features of
DEC types in childhood diarrhea vary with geographical area [6].
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ISSN: 2155-9597
Bacteriology and Parasitology
Yameen et al., J Bacteriol Parasitol 2018, 9:3
DOI: 10.4172/2155-9597.1000339
Research Article Open Access
J Bacteriol Parasitol, an open access journal
ISSN:2155-9597
Volume 9 • Issue 3 • 1000339