Journal of Radioanalytical and Nuclear Chemistry, Vol. 269, No.3 (2006) 579–583
0236–5731/USD 20.00 Akadémiai Kiadó, Budapest
© 2006 Akadémiai Kiadó, Budapest Springer, Dordrecht
Biodistribution of gyroxin using
125
I as radiotracer
J. A. Alves da Silva,
1
* E. Muramoto,
2
M. T. C. P. Ribela,
1
J. R. Rogero,
1
M. A. P. Camillo
1
1
Molecular Biology Center, IPEN/CNEN-SP, P.O.Box 11049, 05422-970, São Paulo, Brazil
2
Radiopharmacy Center, IPEN/CNEN-SP, P.O.Box 11049, 05422-970, São Paulo, Brazil
(Received April 6, 2006)
The use of radiotracers in the research of animal venom has been scarce, although it allows an excellent approach to follow the process of
bioavailability, biodistribution and kinetics of toxins. The purpose of this study was to assess gyroxin action mechanism, transport, compartments
and action sites. This toxin is a thrombin-like and causes the barrel rotation syndrome. The gyroxin was labeled with
125
I and used as a tracer for
the in vivo assay in mice. Blood samples and organs were collected at different time intervals, weighed and analyzed in a gamma-counter. The data
was related with tissues distribution of protease activated receptor (PAR). Biodistribution assay allowed dividing the organs into three groups. The
first one with the organs that followed the blood kinetics, the second with the organs related to metabolisms and elimination, and the third with the
organs in which the gyroxin concentration increased during the observation period.
Introduction
The distribution, metabolism and excretion (DME)
of a strange compound in a living organism depends on
physicals, chemicals and physiological factors
(pharmacokinetics processes). These factors remove and
dilute the compound from its place of entrance in the
biological system, carry it to different tissues, allow its
diffusion or promote its transportation actively through
cells membranes and finally determine its accumulation,
disposition and excretion. The biodistribution rate in the
organ’s tissue is determined by the blood flow perfusion
and by the ability of the compound molecule to pass
through the capillary wall and penetrate into the cells.
1
Gyroxin is a serine protease present in the venom of
Crotalus durissus terrificus, a Brazilian rattlesnake. It
was first described as a neurotoxin causing motor
disturbances called barrel rotation.
2,3
The intravenous
(IV) injection of gyroxin into mice produces temporary
episodes characterized by opisthotonos and rotations
around the long axis of the body: that is the reason why
it is called gyroxin. Gyroxin is also known for its
thrombin-like activity on human fibrinogen, cleaving the
alpha-chain.
4
The fibrin monomers polymerizes into an
abnormal fibrin clot and it is highly susceptible to the
action of fibrinolytic agents.
5,6
The total or partial
incoagulability observed in the blood of patients bitten
by rattlesnakes derives from the fibrinogen
consumption.
7
There is a special interest in gyroxin action on
fibrinogen because of its possible use in thrombotic
disease therapy and as an anticoagulant factor. Some
thrombin-like enzymes isolated from the venom of other
snakes are being used as defibrinogenating agents for a
number of clinical conditions including deep vein
thromboses, myocardial infarction, pulmonary embolus,
acute ischemic stroke and peripheral vascular disease.
5
* E-mail: josilva@ipen.br
As told before, gyroxin is a multifunctional protein
but its action mechanism is not well-known, especially
regarding neurotoxicity. This work aims at investigating
gyroxin action mechanism, transport, compartments and
action sites using
125
I as radiotracer.
The animals’ toxins could be radiolabeled for
biodistribution assays, indirectly or directly:
Indirectly, by labeling the toxin in an already
radiolabeled molecule (e.g., an antibody). This
incorporated radiotracer will show the toxin metabolism
in a living organism during the whole experimental
period.
Directly, by incorporating the radionuclide to the
protein molecule, cofactors, coenzymes or metal, when
one of these constituents is present. The directly way
was used in this study (gyroxin was radiolabeled with
125
I).
Radiolabeling with
125
I has been indicated for
several applications due to its easy and simple
radioiodination system, labeled product stability and
detection facility.
8
Experimental
Gyroxin radioiodination
Gyroxin was obtained from Crotalus durissus
terrificus’ crude venom by affinity and gel filtration
chromatography.
9
It was radioiodinated using mild
conditions at low temperature with chloramine-T
method.
10
The reaction mixture were fractionated in a
Sephadex G100 column (3×50 cm
2
) with 50mM sodium
phosphate buffer pH 7.4 containing 0.1% of bovine
serum albumin as eluent. Fractions of 2.0 ml/tube were
collected and the flow rate was 12 ml/h. Aliquots with
20 μl from each tube were analyzed in a gamma-counter
(Oakfield Instruments, United Kingdown).