22 Testudo Vol. 9, No. 1
Behaviour of juvenile green turtles
(Chelonia mydas) before and after
fibropapillomatosis tumour removal
Jenny Whilde
1*
, Liam Whitmore
2
, Calvin Yang
1
, Catherine B. Eastman
1
,
Rachel Thomas
1
, Devon Rollinson
1
, Brooke Burkhalter
1
, Mark Q.
Martindale
1
and David J. Duffy
1,2
1
The Whitney Laboratory for Marine Bioscience & Sea Turtle Hospital,
University of Florida, 9505 Ocean Shore Blvd., St. Augustine, FL 32080-8610
2
Department of Biological Sciences, School of Natural Sciences, Faculty of
Science and Engineering, University of Limerick, Limerick, Ireland
*Corresponding author: Jenny Whilde, email: whildej@tcd.ie
Introduction
Wild sea turtle populations worldwide, including in Florida, are under
increasing threat from a fatal virulent tumour disease, fibropapillomatosis (FP)
(Herbst 1994; Duffy et al. 2018; Farrell et al. 2018; Duffy & Martindale 2019;
Jones et al. 2016), potentially undermining turtle conservation efforts (Foley et
al. 2005; Jones et al. 2016). Green sea turtles (Chelonia mydas) are the species
most affected by FP (Fig. 1), but it has also been detected in all other sea turtle
species (Herbst 1994). While FP tumours are generally not malignant, their
detrimental consequences can result in susceptibility to other risks, including
predation, parasitism and secondary diseases and infections (Vittecoq et al.
2015). Tumours can be large enough to restrict feeding, swimming, and
reproductive abilities, and cause immunosuppression (Aguirre & Lutz 2004;
Herbst 1994; Jones et al. 2016; Williams Jr et al. 1994). Currently, surgery is
the primary treatment for turtles with FP, but at least 40% of turtles that have
tumours surgically removed experience regrowth (Page-Karjian et al. 2012;
Duffy et al. 2018; Farrell et al. 2018). Fibropapillomatosis is now spreading
to areas where it has not previously been observed (Duarte et al. 2012; Foley
et al. 2007; Page-Karjian et al. 2014; Hirama & Ehrhart 2007; Hargrove et al.
2016; Foley et al. 2015), which means that the number of stranded turtles
requiring treatment for FP is also increasing.
There are still many gaps in our understanding of sea turtles in general,
with ongoing discussion about the most effective conservation strategies
(Hamann et al. 2010). Research on the behaviour of turtles in the wild tends
to focus more broadly on foraging and migration habits (Dodge et al. 2014;
Hochscheid 2014; Heithaus et al. 2002; Godley et al. 2002). One study made
a comparison between the behaviour of tumour-bearing and non-tumour-
© British Chelonia Group + Jenny Whilde, Liam Whitmore,
Calvin Yang, Catherine B. Eastman, Rachel Thomas,
Devon Rollinson, Brooke Burkhalter, Mark Q. Martindale
and David J. Duffy 2019