© CAB International, 2020. Quantitative Genetics, Genomics and Plant Breeding,
140 2nd Edition (ed. M.S. Kang)
Introduction
Genotype–environment interaction (GEI) is an
age-old, universal issue that relates to all living
organisms (Kang, 1998). Genotypes and environ-
ments interact to produce an array of phenotypes.
GEI is the variation caused by the joint effects of
genotypes and environments (Dickerson, 1962).
Baker (1988) defned GEI as the difference be-
tween the phenotypic value and the value ex-
pected from the corresponding genotypic and
environmental values. When responses of two
genotypes to different levels of environmental
stress are compared, an interaction is described
statistically as the failure of the two response
curves to be parallel (Baker, 1988). Recently, de
Leon et al. (2016) defned GEI as the differential
sensitivity of certain genotypes to different en-
vironments. The GEI issue is not only import-
ant in plant-breeding programmes but also in
animal-breeding programmes (Lin and Lin,
1994; Montaldo, 2001; Hayes et al., 2016).
Genotype-by-environment interaction must
be distinguished from phenotypic plasticity.
Schlichting (1986) defned phenotypic plasticity
as the ability of an individual organism to alter
its physiology/morphology in response to changes
in environmental conditions. It describes the
range of phenotypes produced by a single genotype
in different environments (Grogan et al., 2016).
The different phenotypes are referred to as norms
of reaction (Redei, 1982; Kang, 1998). Pheno-
typic plasticity is a useful mechanism in plants,
as being immobile, plants, unlike animals, can-
not move away from a stressful situation but can
adjust their performance through phenotypic
plasticity. For GEI to occur, there must be at least
two genotypes (df = 1) tested in at least two en-
vironments (df = 1), which will yield GEI with 1
df (Kang, 1998). van Eeuwijk et al. (2016) con-
sider the GEI problem as the building of predict-
ive models for genotype-specifc reaction norms.
We should also distinguish between GEI
and genotype–environment correlation (covari-
ance) (GEC). GEC occurs when phenotypic and
environmental effects are not independent.
When additional agronomic inputs are given or
are necessary for certain genotypes to do well, a
genotype-by-environment correlation is created
(Kang, 1998). Interaction or GEI occurs when
the difference between the average phenotypic
value for two genotypes changes in different en-
vironments, but GEC refers to the situation when
particular genotypes tend to be associated with
positive, and others with negative, environmen-
tal effects (Crow, 1986; Doolittle, 1987).
9 Genotype–Environment Interaction
and Stability Analyses: An Update
Manjit S. Kang*
Kansas State University, Manhattan, Kansas, USA
* Email: manjit675264@gmail.com
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