Molecular dynamics simulations with constrained roto-translational motions: Theoretical basis and statistical mechanical consistency A. Amadei, a) G. Chillemi, b) M. A. Ceruso, A. Grottesi, and A. Di Nola Department of Chemistry, University of Rome ‘‘La Sapienza,’’ P.le A. Moro 5, 00185, Rome, Italy Received 29 April 1999; accepted 4 October 1999 From a specific definition of the roto-translational externaland intramolecular internal coordinates, a constrained dynamics algorithm is derived for removing the roto-translational motions during molecular dynamics simulations, within the leap-frog integration scheme. In the paper the theoretical basis of this new method and its statistical mechanical consistency are reported, together with two applications. © 2000 American Institute of Physics. S0021-96060050201-3 I. INTRODUCTION Often in molecular simulations the interest is focused on the structural properties of a molecule with internal degrees of freedom. For simulations of a single molecule in vacuum ideal gas conditionas well as for simulations of a solute molecule in its solvent infinite dilution condition, the roto- translational motions are in general uninteresting, while the behavior of the internal coordinates can be very important for many different studies. In particular with large organic molecules, and especially biomacromolecules, simulations are usually performed to obtain a detailed investigation of the conformational fluctuations, which can be studied only after removing the overall translation and rotation of the molecule. In these cases the ensemble of molecular configu- rations obtained from the simulation is normally manipulated to remove these roto-translational motions. In general the roto-translation is eliminated by over-imposing the center of mass of the actual configuration with that of a reference one, and then least square fitting the atomic displacements be- tween the two structures rotating the actual structure around its center of mass. 1 This procedure, although usually efficient and widely used, has one disadvantage. Its implicit definition of external and internal coordinates is rather complicated, especially for the definition of conjugated momenta, and hence it is difficult to use this approach for theoretical me- chanics or rigorous statistical mechanical studies, as well as to derive ideal constraint forces to stop the molecular roto- translational motions directly during the simulation. It should be considered that a rigorous method to constrain the roto- translational motions during a simulation can be advanta- geous in the following cases: 1For large and flexible molecules the removal of the dy- namical coupling between the internal motions and mo- lecular roto-translations can shorten the system’s relax- ation time and hence provide a better configurational sampling for the internal coordinates in simulations of usual time lengths. 2Simulations of a molecule in vacuum, with either a usual force field or using a mean field, are usually performed at zero angular momentum and this constraint can alter the statistical mechanical consistency of the simulation. On the contrary when ideal holonomic constraints are used to stop the roto-translational motions, and the an- gular momentum is not fixed anymore, the simulation can provide the exact statistical mechanics of the system. 3For simulations of large nonspherical molecules e.g., proteinsin water, with the presence of the roto- translational constraints we could use a simulation box shaped on molecular geometry, reducing significantly the number of necessary water molecules. The effect of large solute rotations is negligible for short length simu- lations hundreds of picosecondsbut becomes relevant for longer time simulations. In fact, from the nanosec- onds range the solute has enough time to rotate signifi- cantly and hence, without using a cubic simulation box, to interact directly with its periodic images. 4For the calculation of free energy differences due to changes of the roto-translational configuration for inter- acting molecules or for a molecule interacting with an external field e.g., molecular dockingthe simulations with the roto-translational constraints could be extremely efficient. In this paper we show that it is possible to use a defini- tion of internal and external coordinates which is very suited for theoretical derivations and that allows direct simulation of only the molecular internal degrees of freedom. In fact, this definition of the molecular coordinates standard in ana- lytical mechanicsallows the use of ideal constraint forces to stop instantaneously the roto-translational motions during the simulation. It is well known from theoretical mechanics 2,3 that the use of ideal holonomic constraints in a Hamiltonian system still provides Hamiltonian dynamics in the con- strained phase space constraint subspace, and so a con- strained Hamiltonian system can still be described by the microcanonical ensemble. We will show that more often in general the holonomic constraints do not alter the basic type of dynamics in the case of the usual molecular dynamics MDequations of motion involving a frictional term, and a Author to whom correspondence should be addressed; electronic mail: amadei@seurat.chem.uniroma1.it b Inter-University Computing Consortium CASPUR, University of Rome ‘‘La Sapienza,’’ P.le A. Moro 5, 00185, Rome, Italy. JOURNAL OF CHEMICAL PHYSICS VOLUME 112, NUMBER 1 1 JANUARY 2000 9 0021-9606/2000/112(1)/9/15/$17.00 © 2000 American Institute of Physics Downloaded 13 Jan 2003 to 151.100.52.54. Redistribution subject to AIP license or copyright, see http://ojps.aip.org/jcpo/jcpcr.jsp