Vol.:(0123456789) 1 3
Journal of Endocrinological Investigation
https://doi.org/10.1007/s40618-020-01328-z
ORIGINAL ARTICLE
Chronic stress inhibits hypothalamus–pituitary–thyroid axis
and brown adipose tissue responses to acute cold exposure in male
rats
A. Castillo‑Campos
1
· A. Gutiérrez‑Mata
1
· J.‑L. Charli
1
· P. Joseph‑Bravo
1
Received: 26 December 2019 / Accepted: 5 June 2020
© Italian Society of Endocrinology (SIE) 2020
Abstract
Purpose Cold exposure activates the hypothalamus–pituitary–thyroid (HPT) axis, response blunted by previous acute stress
or corticosterone administration. Chronic stressors can decrease serum T3 concentration, and thyrotropin-releasing hormone
(Trh) expression in the paraventricular nucleus (PVN), but impact on the response to cold is unknown; this was studied in rats
submitted to daily repeated restraint (rRes) that causes habituation of hypothalamus–pituitary–adrenal (HPA) axis response,
or to chronic variable stress (CVS) that causes sensitization and hyperreactivity.
Methods Wistar male adult rats were submitted to rRes 30 min/day, or to CVS twice a day, for 15 days. On day 16, rats were
exposed 1 h to either 5 or 21 °C. Parameters of HPT and HPA axes activity and of brown adipose tissue (BAT) cold response
were measured; gene expression in PVN and BAT, by RT-PCR; serum hormone concentration by radioimmunoassay or
ELISA.
Results Compared to naïve animals, Crh and corticosterone concentrations were attenuated at the end of rRes, but increased
at the end of CVS treatments. Cold exposure increased mRNA levels of Crh, Trh, and serum concentration of thyrotropin in
naïve, but not in rRes or CVS rats; corticosterone increased in all groups. Cold induced expression of thermogenic genes in
BAT (Dio2 and Ucp1) in naïve but not in stressed rats; Adrb3 expression was diferentially regulated.
Conclusion Both types of chronic stress blunted HPT and BAT responses to cold. Long-term stress efects on noradrenergic
and/or hormonal signaling are likely responsible for HPT dysfunction and not the type of chronic stressor.
Keywords Thyrotropin-releasing hormone · Brown adipose tissue · Hypothalamus–pituitary–thyroid · Hypothalamus–
pituitary–adrenal · Restraint · Chronic variable stress · Cold stress
Abbreviations
ACTH Adrenocorticotropin
Adrb3 β3 Adrenergic receptor gene
BAT Brown adipose tissue
BWg Body weight gain
cAMP Cyclic adenosine monophosphate
Cort Corticosterone
Cpf Control pair-fed
Crh Corticotropin-releasing hormone gene
CRH Corticotropin-releasing hormone peptide
CVS Chronic variable stress
Dio2 Deiodinase 2 gene
DIO2 Deiodinase 2
EPM Elevated plus maze
Gr Glucocorticoid receptor gene (Nr3c1: nuclear
receptor subfamily 3 group C member gene)
HPA Hypothalamus–pituitary–adrenal axis
HPT Hypothalamus–pituitary–thyroid axis
N Naïve
OFT Open feld test
pCREB Phosphorylated cAMP response element-bind-
ing protein
PKA Protein kinase A
PKAc Catalytic subunit of PKA
A. Castillo-Campos and A. Mata made equivalent contributions.
Electronic supplementary material The online version of this
article (https://doi.org/10.1007/s40618-020-01328-z) contains
supplementary material, which is available to authorized users.
* P. Joseph-Bravo
joseph@ibt.unam.mx
1
Departamento de Genética del Desarrollo y Fisiología
Molecular, Instituto de Biotecnología, Universidad Nacional
Autónoma de México (UNAM), Cuernavaca, México